Diet selection in pigs: choices made by growing pigs given foods of different protein concentrations

1990 ◽  
Vol 51 (1) ◽  
pp. 189-199 ◽  
Author(s):  
I. Kyriazakis ◽  
G. C. Emmans ◽  
C. T. Whittemore
Keyword(s):  
Growth Rate ◽  
Food Intake ◽  
Protein Concentration ◽  
Live Weight ◽  
Diet Selection ◽  
Growing Pigs ◽  
Growth Potential ◽  
Feeding Method ◽  
Total Food Intake ◽  
Total Food

ABSTRACTTo test the proposition that growing pigs, when given a choice between two foods, are able to select a diet that meets their requirements, and to investigate the rules of diet selection, four foods (L, A, B and H) with similar energy yields, but different concentrations of crude protein (CP) (125, 174, 213 and 267 g CP per kg fresh food respectively) were formulated. The four foods were offeredad libitumeither singly, or as a two-way choice using all the six possible pairs, to 40 individually caged pigs from 12 to 30 kg live weight. On the single foods the rate of food intake fell from 1001 to 971 to 961 to 868 (s.e.d. 40) g/day (F < 0·05) as the protein concentration of the foods increased from L to H; the growth rate followed an opposite trend (492, 627, 743 and 693 (s.e.d. 31) g/day respectively;P< 0·01). When the pigs had to select between two foods limiting in protein (L and A) the less limiting one was preferred (710 (s.e. 200) g A per kg total food intake; the protein concentration of the selected diet was 160 (s.e. 10) g CP per kg). On the choice between B and H (a choice between a food with protein concentration close to requirements and a food with protein excess) the lower food was markedly preferred (928 (s.e. 4) g B per kg total food intake; the protein concentration of the selected diet was 218 (s.e. 1) g CP per kg). When the animals were given a choice between two foods, a combination of which was non-limiting (pairs LB, LH, AB and AH), the protein concentrations of the selected diets were not different between treatments (208, 204, 202 and 205 (s.e.d. 13) g CP per kg respectively) and they also declined systematically with time and weight. The growth rate of the animals on these pairs were 752, 768, 769 and 763 (s.e.d. 54) g/day (P > 0·05), which were not significantly different from the highest growth rate achieved on a single food. The results suggest that pigs, when given a choice between a suitable pair of foods, are able to choose a balanced diet and to change its composition to reflect their changing requirements. The choice-feeding method may well be useful as an effective and economic way of estimating and meeting requirements, and of measuring the growth potential of pigs.

Technical review of the energy and protein requirements of growing pigs: food intake

2001 ◽  
Vol 73 (1) ◽  
pp. 3-17 ◽  
Author(s):  
C. T. Whittemore ◽  
D. M. Green ◽  
P. W. Knap
Keyword(s):  
Food Intake ◽  
Stocking Density ◽  
Live Weight ◽  
Growth Period ◽  
Growing Pigs ◽  
Potential Growth ◽  
Total Food Intake ◽  
Total Food ◽  
Minimum Number ◽  
Reasonable Prediction

AbstractFood intake in pigs is highly variable across different production circumstances. This report concludes from a critical review of published observations that it was unrealistic to expect from the scientific literature purporting to express nutrient requirement any reasonable prediction of the particular food intake of groups of pigs. None the less, such knowledge is essential for the practical purposes of their day-to-day nutrition. The literature does however yield general principles from which may be derived: (a) the likely forms (but not the parameter values) of intake functions relating food intake to pig live weight; and (b) the likely factors involved in the modulation of food intake at any given live weight. Using these principles two methods for determining on-farm food intake from the use of simple and available records were proposed. The first requires knowledge only of start and final weight, the time elapsed, and total food intake: it involves two steps, the determination of a suitable growth curve followed by the fitting of a suitable food intake curve. The second method is appropriate in the absence of information on total food intake, and requires a minimum number of spot measurements through the growth period. Different functions were tested for the curve of best fit. As a further benefit it appeared that models could be constructed from the information presented that would speculate for diagnostic purposes upon the likely modulators of food intake. Such models could explore the constraints of gut capacity, the energetic requirements of maintenance and potential growth, the influence of excessive or inadequate environmental temperature, the quality of housing and stocking density.

Tests of two theories of food intake using growing pigs 2. The effect of a period of reduced growth rate on the subsequent intake of foods of differing bulk content

2001 ◽  
Vol 72 (2) ◽  
pp. 361-373 ◽  
Author(s):  
E.C. Whittemore ◽  
G.C. Emmans ◽  
B.J. Tolkamp ◽  
I. Kyriazakis
Keyword(s):  
Growth Rate ◽  
Food Intake ◽  
Sugar Beet ◽  
Live Weight ◽  
Growing Pigs ◽  
Sugar Beet Pulp ◽  
Beet Pulp ◽  
Bulk Content ◽  
High Bulk ◽  
Subsequent Intake

AbstractThe effect of a period of feeding on a high bulk food, upon the subsequent intake of foods of differing bulk content, was investigated in two experiments of the same design. The intention was to provide a severe test of the two current conceptual frameworks available for the prediction and understanding of food intake. In each experiment 40 male Manor Meishan pigs were randomly allocated to one of four treatment groups at weaning. Each experiment was split into two periods, P1 (12 to 18 kg) and P2 (18 to 32 kg). The treatments, all with ad libitum feeding, were: a control food (C) given throughout (treatment CC); a medium bulk food (M) given throughout (treatment MM); a high bulk food (H) given in P1 and then C in P2 (treatment HC); H given in P1 and M in P2 (treatment HM). C was based on micronized wheat with 13·4 MJ digestible energy and 243 g crude protein per kg fresh food. In experiment 1 M contained 350 g/kg and H 560 g/kg of unmolassed sugar-beet pulp and in experiment 2 M contained 500 g/kg and H 700 g/kg of unmolassed sugar-beet pulp. Framework 1 predicted that food intake on the medium bulk food (M) would not be increased, whereas framework 2 predicted that intake on M would be increased after a period of feeding on H, compared with when M was offered continuously.In P1, both food intake (P < 0·01) and growth (P < 0·001) were severely limited on H compared with C. In experiment 1 growth was limited on M compared with C during the first 7 days of P1 (P < 0·01) only. In experiment 2 intake (P < 0·001) and growth (P < 0·001) on M were limited throughout P1, compared with C but not thereafter. Therefore, in neither experiment did M cause a lower growth rate than C from 18 to 32 kg. In experiment 1 there was full adaptation to M after about 10 days from 12 kg. In experiment 2 adaptation was complete by the end of the first 7 days from 18 kg.In P2, food intake (P < 0·001) and live-weight gain (P < 0·05 and P < 0·001 in experiments 1 and 2, respectively) were increased on HC compared with CC. By the last 7 days of P2 intake was still higher (P < 0·01) but growth rate was no longer different to CC. Intake and gain were increased in P2 on HM compared with MM but, in general, these differences were small and not significant. In the first 7 days of P2, in experiment 1 pigs on HM had higher intakes (P < 0·001) and gains (P < 0·05) than those on MM, but in experiment 2 only intake was higher (P < 0·01) with no difference in gain. By the last 7 days of P2 there was no difference in either intake or gain between these two groups in either experiment. Pigs on HC increased intake by more than those on HM. There was, therefore, a significant interaction for food intake (P < 0·05, in experiment 1 and P < 0·001, in experiment 2) between prior and present food.The unexpected failure of either M food to limit growth throughout the experimental period meant that the results of these experiments could not be used as a strong test to reject either one of the frameworks. However, the ability of the pigs to compensate on M was less than that on C. The data provide some evidence that under conditions of compensation foods such as M may be limiting. This is in closer agreement with the framework that predicted that consumption of a limiting food will not increase after a period of feeding on a high bulk food (framework 1).

The effect of stocking density on the responses of growing pigs to dietary lysine

2001 ◽  
Vol 73 (3) ◽  
pp. 459-469 ◽  
Author(s):  
N.S. Ferguson ◽  
G. Lavers ◽  
R.M. Gous
Keyword(s):  
Growth Rate ◽  
Food Intake ◽  
Stocking Density ◽  
Live Weight ◽  
Growing Pigs ◽  
Lysine Content ◽  
Large White ◽  
Daily Growth ◽  
Body Protein ◽  
Floor Space

AbstractAn experiment was conducted to measure the effects of stocking density (increased number of pigs per pen) on lysine requirements of pigs grown from 25 to 60 kg live weight. Two hundred and sixty-four female Large White ✕ Landrace pigs were assigned at 25 kg to one of four dietary lysine treatments (13·3 (L1); 11·4 (L2); 9·5 (L3) and 7·6 (L4) g/kg) and either seven or 13 pigs per pen (or 1 0 and 0·5 m2 per pig, respectively). An additional treatment of one pig per pen (20 m2 per pig) was included to compare the responses of solitary- versus group-penned pigs. Animals were given ad libitum access to dietary treatments from a mean pen starting weight of 261 (s.e. 0·35) kg to a mean pen finishing weight of 63·4 (s.e. 0·61) kg live weight. There were no significant interactions between dietary lysine content and floor space per pig on food intake (FI), average daily growth rate (ADG), the amount of food per unit of gain (FCR) and the rate of protein retention (PR). Significant interactions were evident for body composition and the rate of lipid retention (LR). Over the weight range 25 to 40 kg there were significant differences in FI (P < 0·05) and FCR (P < 0·001) between dietary lysine treatments but most of these differences had disappeared over the 40 to 60 kg live weight. Individually penned animals had significantly higher (P < 0·05) FI and ADG than group-penned animals. However, there were no differences between seven and 13 pig per pen treatments. Stocking density had no effect on LR or body protein content but did cause a significant reduction in PR (P < 0·001) and an increase in body lipid content (P < 0·05) as the number of pigs per pen increased from seven to 13. Lysine requirements (expressed in g/day) therefore could be seen to be reduced with increasing stocking density. However, as lysine intake was reduced in group-penned animals, the reduced daily requirement does not necessarily warrant a reduction in the lysine content of the food. Feeding according to the requirements for maximum PR will still produce the best carcass and growth performance irrespective of the group size. The improvement in PR associated with higher dietary nutrient levels did not completely offset the adverse physiological effects of higher stocking density but may partly counteract the effect of reduced lysine intake. However, there were indications that feeding crowded pigs a lower dietary lysine concentration may not further reduce the already diminished protein (lysine) growth rate. An additional experiment was performed to test whether the number of feeder bins may have constrained food intake and therefore growth in group-penned animals. The results of this experiment showed that the number of bins had no significant effect on FI, ADG and FCR in group-penned pigs, and therefore a single feeder bin was not considered a constraining factor in pigs housed with limited floor space.

The effect of a methane-suppressing compound, trichloroethyl adipate, on rumen fermentation and the growth of sheep

1977 ◽  
Vol 24 (2) ◽  
pp. 169-181 ◽  
Author(s):  
J. L. Clapperton
Keyword(s):  
Weight Gain ◽  
Food Intake ◽  
Live Weight ◽  
Rumen Fermentation ◽  
Total Food Intake ◽  
Total Food ◽  
Four Levels ◽  
Lamb Growth

SUMMARY1. Three experiments have been carried out to study the effect of adding trichloroethyl adipate, a compound known to prevent the production of methane bothin vitroandin vivo, to the diet of sheep on fermentation in the rumen and on the growth of lambs. In Experiment 1 the additive was incorporated at various levels for 6 weeks, in Experiment 2 it was given at one of four levels for 12 weeks and in Experiment 3 it was included in the food of growing lambs for 15 weeks.2. Initially, in Experiments 1 and 2, the additive caused a large reduction in methane production and in the molar proportion of acetic acid in the rumen liquor with a corresponding increase in propionic acid. These effects were much reduced after the first 2 to 3 weeks of administration.3. The total food intake of the animals receiving the additive in Experiment 3 was 4% less than that of the control animals. This difference was negligible at the start of the experiment but increased throughout the feeding period.4. On average, the relative live-weight gain and the changes in the weights of carcass, viscera and empty body weight of the animals receiving the additive were also 4% less than those of the control animals.5. The observed differences in weight gain resulted from the differences in food intake and the additive had no beneficial effect upon lamb growth. This lack of effect on growth was due to rapid adaptation by the rumen fermentation to the presence of the additive.

Effect of food intake capacity on genotype by feeding regimen interactions in growing pigs

1990 ◽  
Vol 50 (2) ◽  
pp. 343-351 ◽  
Author(s):  
E. Kanis
Keyword(s):  
Growth Rate ◽  
Food Intake ◽  
Live Weight ◽  
Tissue Growth ◽  
Growing Pigs ◽  
Fatty Tissue ◽  
Restricted Feeding ◽  
Production Traits ◽  
Feeding Regimen ◽  
Ad Libitum

ABSTRACTFrom each of 159 litters, between one and three pigs were fed ad libitum and between one and four pigs were fed at several restricted feeding levels from about 27 to 107 kg live weight. Effects of litter by feeding regimen (ad libitumv. restricted feeding) interactions (L × F) and sex by feeding regimen interactions (S × F) on production traits were investigated. In experimental groups where restrictedly fed animals received an amount of food based on their estimated voluntary food intake capacity (FIC), interactions were non-significant. In groups where restricted feeding was independent of FIC, significant L × F or S × F interactions were found for daily food intake (FI), daily gain, lean tissue growth rate and fatty tissue growth rate, but not for body composition and food conversion traits.Because of between-animal variation in FIC, restricted feeding according to a fixed scale restricted intake of some animals to a higher degree than others and degree of food intake restriction (DFR) was calculated from actual FI and FIC for animals fed at a restricted level. After correction of FI and production traits of restrictedly fed animals for differences in DFR, L × F and S × F interactions were not significant.It was concluded that the poor relationships often found between test-station results of boars and results of their progeny in practical environments may be caused, to an important extent, by differences in DFR in each environment. To overcome these genotype by environment interactions it is recommended that animals be fed ad libitum in both test and commercial environments.

Reproductive performance, body weight and body condition of breeding sows with differing body fatness at parturition, differing nutrition during lactation, and differing litter size

1989 ◽  
Vol 48 (1) ◽  
pp. 181-201 ◽  
Author(s):  
H. Yang ◽  
P. R. Eastham ◽  
P. Phillips ◽  
C. T. Whittemore
Keyword(s):  
Food Intake ◽  
Litter Size ◽  
Post Partum ◽  
Live Weight ◽  
Size Change ◽  
Large White ◽  
Feeding Level ◽  
Total Food Intake ◽  
Total Food ◽  
High Level

ABSTRACTOne hundred and two Large White × Landrace Fl hybrid sows were allocated over four parities in a factorial design to two levels of target P2 backfat thickness at parturition (20 mm, F v. 12 mm, T), two levels of lactation feeding {ad libitum, H v. 3 kg/day, L), and two sizes of sucking litter (six v. 10). Sows attained 13 mm P2 when first mated at 126 kg live weight. Fatness (P2, mm) at weaning was significantly influenced by target fatness at parturition (14·2 v. 9·3), lactation feeding level (13·7 v. 10·0), and litter size (12·7 v. 11·0). Changes in backfat (P2, mm) during 28-day lactation were significantly influenced by target fatness at parturition (—5·0 v. —2·5), lactation feeding (-2 0 v. —5·4), and litter size (—2·9 v. -4·6). Sow live weight (kg) at weaning was significantly influenced by target fatness at parturition (211 v. 192), lactation feeding (218 v. 186), and litter size (208 v. 196). Changes in live weight (kg) during 28-day lactation were significantly influenced by target fatness at parturition (—26 v. — 12), lactation feeding (—5 v. —31), and litter size (—12 v. —25). With multiparous sows only, total food intake during 28-day lactation was negatively related to total food intake in pregnancy. Change in backfat (P2, mm) during 28-day lactation = -0·28 - 0·27 P2 at parturition + 0·04 lactation food intake — 0·50 litter size. Change in live weight (kg) during 28-day lactation = -3·8 — 0·15 live weight post partum + 0·36 lactation food intake — 3·3 litter size. Sows with target fat levels of 20 mm P2 at parturition had better food conversion efficiencies than sows with target fat levels of 12 mm. Target fatness at parturition, and especially lactation food intake, but not litter size, significantly influenced the interval (days) from weaning to oestrus in parity 1 (9·1 v. 14·2 and 7·8 v. 15·3, but 11·6 v. 11·5), while n i subsequent parities only litter size influenced the interval (days) from weaning to oestrus (6·0 v. 8·0). Birth weight (kg) of piglets was influenced only marginally by target fatness at parturition (1·4 v. 1·2) in parity 1, and not by the other factors, or in subsequent parities. Piglet growth rate was affected by both target fatness at parturition and litter size, but by lactation feeding level only in the last week of lactation. The relationship between fatness at weaning (mm) and the weaning to oestrus interval (days) for primiparous sows can be expressed as 26·6 — 1·28 P2. High level feeding in lactation imparted production and efficiency benefit in both primiparous and multiparous sows, while pregnancy feeding to a target of 20 mm rather than 12 mm at parturition was of benefit for primiparous sows.

The effects of dietary fibre, source of fat and dietary energy concentration on the voluntary food intake and performance of growing pigs

1986 ◽  
Vol 43 (2) ◽  
pp. 327-333 ◽  
Author(s):  
R. G. Campbell ◽  
M. R. Taverner
Keyword(s):  
Growth Rate ◽  
Food Intake ◽  
Energy Intake ◽  
Live Weight ◽  
Growing Pigs ◽  
Energy Concentration ◽  
Dietary Energy ◽  
Crude Fibre ◽  
Acid Detergent Fibre ◽  
Voluntary Food Intake

ABSTRACTNinety entire male pigs were used in two experiments to investigate the effects of energy concentration on the performance of growing pigs given diets containing relatively high or low levels of fibre. A blended fat product was also compared with tallow as a source of dietary energy for pigs in the first experiment.In experiment 1, 60 pigs growing between 20 and 45 kg live weight were given 10 diets containing approximately 120 g acid-detergent fibre per kg (80 g crude fibre per kg), and 10, 30, 50, 75 or 100 g/kg of either tallow or the blended fat product.Although the source of fat had no significant effect on food intake or growth performance, voluntary energy intake and growth rate increased with increase in the level of fat added to the diet and with concomitant increase in dietary digestible energy (DE) concentration up to 50 g/kg and 13·2 MJ/kg respectively. Further increase in the level of fat added to the diet and in DE concentration resulted in a decline in voluntary food intake, whilst energy intake and growth rate remained relatively constant at 27 MJ/day and 720 g/day respectively.In experiment 2, 30 pigs were used to investigate the animals' response to five levels of dietary DE concentration (11·8 to 15·1 MJ DE per kg) between 20 and 50 kg live weight. The highest level of acid-detergent fibre in the diets was 62 g/kg (50 g crude fibre per kg) and this declined with increase in energy concentration to 30 g/kg (25 g crude fibre per kg) in the diet of highest energy concentration.Voluntary food intake was not significantly affected by dietary energy concentration, and voluntary energy intake and growth rate increased linearly with increase in dietary energy concentration up to 14·5 MJ DE per kg. Maximum energy intake (31 MJ DE per day) and growth rate (905 g/day) in experiment 2 was proportionately 0·15 and 0·26 higher than that achieved by pigs given the higher fibre diets in experiment 1.The results for voluntary food intake and growth rate suggested that the pig's demand for energy and consequently its response to dietary energy concentration was affected by dietary fibre concentration.

Diet selection of sheep: sodium bicarbonate, but not the offering of hay, modifies the effect of urea on diet selection

2002 ◽  
Vol 74 (2) ◽  
pp. 357-367 ◽  
Author(s):  
S. M. James ◽  
I. Kyriazakis ◽  
G. C. Emmans ◽  
B. J. Tolkamp
Keyword(s):  
Sodium Bicarbonate ◽  
Live Weight ◽  
Diet Selection ◽  
Metabolizable Energy ◽  
Fresh Matter ◽  
Total Food Intake ◽  
Period 2 ◽  
Control Food ◽  
Selection Of

AbstractThe hypotheses tested were that the expected preference of sheep for a food with adequate rumen degradable protein (RDP) supplemented with urea would be reduced both by the addition of a buffer (sodium bicarbonate (SB)) and by offering ad libitum access to hay. A control food (C), calculated to be adequate in its ratio of effective RDP to fermentable metabolizable energy (fME), was formulated. Other foods were made by adding 12·5 (U1) or 25 (U2) g urea per kg fresh matter (FM) to C and 20 g SB per kg FM to C, U1and U2. The acid buffering capacity (ABC) of each food was measured in vitro. The experiment consisted of two successive periods, each of 4 weeks. Ninety-eight female, Texel ✕ Greyface sheep were randomly allocated to 14 groups each with seven animals. Groups 1 to 6 were offered one of: C, U2, C + SB, U2+ SB, C with hay or U2with hay throughout the experiment. Groups 7 to 10 were offered the choices of C v. U1or C v. U2, either with or without hay in a change-over design; animals that received hay during period 1 (groups 8 and 10) did not do so during period 2 and vice versa (groups 7 and 9). Groups 11 to 14 (no. = 7) were offered the choices of C v. U1or C v. U2, either with or without SB supplemented to both foods, in a change-over design. Adding either urea, or SB, or both to C had no effects on intake or live-weight gain when offered alone. Both supplements significantly (P 0·001) increased the ABC of food C. Throughout the experiment hay consumption was very low (overall mean: 23 (s.e. 2·5) g hay per sheep day). Offering hay caused no change in the preference for the urea-supplemented foods. Sheep offered the choices C v. U1or C v. U2, with neither hay nor SB, selected 0.466 (s.e. 0·036) and 0.588 (s.e. 0·025) kg/kg total food intake (TFI) of U1and U2respectively. The proportions of the urea-supplemented foods were significantly reduced (P 0.01) by SB supplementation: to 0.348 (s.e.0·045) and 0·406 (s.e.0·059) kg/kg TFI of U1and U2respectively. The effect of SB addition on the diet selection of sheep could be due to its buffering properties. When SB is added to both foods the need for urea to be used as a buffer is reduced with a consequent decrease in the proportion selected as the urea-supplemented food. Effects of diet on buffering may override other diet selection objectives, such as the avoidance of an excess intake of RDP.

Influence of energy and protein concentration in the diet on the performance of growing pigs 2. Differing nutrient density at a constant energy: protein ratio

1972 ◽  
Vol 14 (1) ◽  
pp. 47-55 ◽  
Author(s):  
G. A. Lodge ◽  
M. E. Cundy ◽  
R. Cooke ◽  
D. Lewis
Keyword(s):  
Growth Rate ◽  
Live Weight ◽  
Fat Content ◽  
Growing Pigs ◽  
Feed Conversion ◽  
Large White ◽  
Protein Ratio ◽  
Treatment Groups ◽  
Nutrient Density ◽  
Amino Acid Balance

SUMMARYForty-eight gilts by Landrace sires on Large White × Landrace females were randomly allocated to eight pens and within pens to six treatment groups involving three diets and two levels of feeding from 23 to 59 kg live weight. All diets were formulated to have approximately the same ratio of digestible energy to crude protein (160 kcal DE/unit % CP) but different energy and protein concentrations: (A) 3500 kcal/kg DE and 21 % CP, (B) 3150 kcal/kg DE and 19% CP, and (C) 2800 kcal/kg DE and 17% CP. Amino acid balance was maintained relatively constant with synthetic lysine, methionine and tryptophan. The levels of feeding were such that the lower level of diet A allowed an intake of energy and protein similar to the higher level of diet B, and the lower level of B was similar to the higher level of C.On the lower level of feeding, growth rate, efficiency of feed conversion and carcass fat content increased linearly with each increment in nutrient concentration; on the higher level of feeding growth rate and EFC increased from diet C to B but not from B to A, whereas carcass fat content increased linearly with diet from the lowest to the highest concentration. There was a non-significant tendency for the higher density diets at a similar level of nutrient intake to give better EFC and fatter carcasses than the lower density diets.

A note on the response of pigs weaned at 28 days to dietary protein

1986 ◽  
Vol 42 (3) ◽  
pp. 427-429 ◽  
Author(s):  
R. G. Campbell ◽  
M. R. Taverner
Keyword(s):  
Growth Rate ◽  
Food Intake ◽  
Dietary Protein ◽  
Live Weight ◽  
Gain Ratio ◽  
Digestible Energy ◽  
Voluntary Food Intake

ABSTRACTThirty-six piglets were used to investigate the effect of six concentrations of dietary protein ranging from 155 to 235 g/kg, and corresponding dietary lysine concentrations from 10·1 to 15·4 g/kg, on the performance of pigs weaned at 28 days of age and growing between 7·5 and 20 kg live weight. Voluntary food intake was not significantly affected by dietary protein, and growth rate increased with increase in dietary protein and lysine up to 167 and 10·9 g/kg respectively (0·75 g lysine per MJ digestible energy (DE)). Food: gain ratio improved significantly with each increase in dietary protein and lysine up to 177 and 11·6 g/kg (0·79 g lysine per MJ DE) respectively.

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