Coleopterous egg deposition on alder leaves from the Klondike Mountain Formation (Middle Eocene), northeastern Washington

1991 ◽  
Vol 65 (2) ◽  
pp. 334-335 ◽  
Author(s):  
Standley E. Lewis ◽  
Mark A. Carroll

Flea beetle (Chrysomelidae) egg deposition has been found on three impressions of alder (Alnus parvifolia) leaves collected at a roadcut in Republic, Washington. These fossils were discovered by Wes Wehr (University of Washington, Burke Museum) during investigations of fossil plants from the Republic, Washington, area. These impressions represent a yet to be determined species, belonging to the genus Altica GeofFroy (1762). They represent the first member of this genus to be described from the Eocene of North America. The fossils were found in lacustrine rocks from the lower part of the Klondike Mountain Formation. Figure 1 represents the Republic fossil locality and the distribution of this Formation. A brief description of the geologic history of this region can be found in Wolfe and Wehr (1987). The Klondike Mountain Formation has a radiometric age that ranges from 42.3 ± 2.0 to 50.3 ± 1.7 m.y. (Pearson and Obradovich, 1977).

1983 ◽  
Vol 120 (1) ◽  
pp. 51-58 ◽  
Author(s):  
A. J. Boucot ◽  
C. H. C. Brunton ◽  
J. N. Theron

SummaryThe Devonian brachiopod Tropidoleptus is recognized for the first time in South Africa. It is present in the lower part of the Witteberg Group at four widely separated localities. Data regarding the stratigraphical range of the genus elsewhere, combined with information on recently described fossil plants and vertebrates from underlying strata of the upper Bokkeveld Group, suggest that a Frasnian or even Givetian age is reasonable for the lower part of the Witteberg Group. The recognition of Tropidoleptus in a shallow water, near-shore, molluscan association, at the top of the South African marine Devonian sequence, is similar to its occurrence in Bolivia, and suggests a common Malvinokaffric Realm history of shallowing, prior to later Devonian or early Carboniferous non-marine sedimentation. It is noteworthy that Tropidoleptus is now known to occur in ecologically suitable environments around the Atlantic, but is absent from these same environments in Asia and Australia. Tropidoleptus is an excellent example of dispersal in geological time — first appearing in northern Europe and Nova Scotia, then elsewhere in eastern North America and North Africa, followed by South America and South Africa, while continuing in North America.


2020 ◽  
Vol 55 (2) ◽  
pp. 91-129
Author(s):  
Jason A. Lillegraven

ABSTRACT The geologic history of Wyoming’s Hanna Basin is still being written. Surprisingly, here appeared an opportunity to share insights from previously accomplished work with that conducted anew by other scholars. The area of study was in the southeastern quadrant of Wyoming, which exhibits the state’s most complex history with respect to the Laramide orogeny. Especially important for present purposes were the tectonic conditions of the late Paleocene and earliest Eocene, recorded within the Hanna Formation. Of central focus is the 2020 publication by Dechesne and her six co-authors. Geographically, the landscape they covered was a thin, synclinal slice of the northeastern margin of the Hanna Basin. Key goals for the present publication have been to illustrate positive linkages and to highlight discrepancies between Dechesne et al. (2020) and relevant prior geological work. A concern that permeates all facets of this approach is the ability to verify viability of brand-new geologic descriptions, data, and resulting conclusions. Essential graphical elements were introduced first into this present publication. Once that package of background information was available, more focused analyses were rigorously pursued on diverse issues within the Dechesne et al. (2020) publication. Dechesne’s team presented a significantly modified but adequately defended approximation of the Paleocene–Eocene boundary. Data from fossil plants (macro- and palynofloras), continental mollusks, and bulk organic-carbon isotopes all agree within one measured section (of five sections studied) with an approximated Paleocene–Eocene boundary along with a ‘carbon isotope excursion’ (CIE). Strength of available evidence seems questionable, however, in that the inordinately high variability in bulk organic carbon (characteristic of a CIE) has been demonstrated only in the Hanna Draw Section. Although fluvial, paludal, and lacustrine facies are considered in several contexts, in no sense does the publication’s organizational form provide a ‘detailed stratigraphic framework.’ One zircon-based U–Pb depositional date (54.42 ± 0.27 Ma) came from this study that matched early Wasatchian time. Participants in the Dechesne et al. (2020) project are to be commended in that their resulting paper ranged broadly across the geologic setting, stratigraphy, paleocurrents, paleobotany, continental mollusks, zircon geochronology, associated lithofacies, and paleogeography. Despite that breadth, there exists a plethora of unexpected and wholly avoidable inconsistencies, strong contradictions within what should be homogeneous datasets, and seemingly inexplicable omissions of obviously necessary and sometimes clearly existing but unutilized data, one must question the reliability of much of the information presented in their paper.


PeerJ ◽  
2016 ◽  
Vol 4 ◽  
pp. e2639 ◽  
Author(s):  
Matthew R. Borths ◽  
Patricia A. Holroyd ◽  
Erik R. Seiffert

Hyaenodonta is a diverse, extinct group of carnivorous mammals that included weasel- to rhinoceros-sized species. The oldest-known hyaenodont fossils are from the middle Paleocene of North Africa and the antiquity of the group in Afro-Arabia led to the hypothesis that it originated there and dispersed to Asia, Europe, and North America. Here we describe two new hyaenodont species based on the oldest hyaenodont cranial specimens known from Afro-Arabia. The material was collected from the latest Eocene Locality 41 (L-41, ∼34 Ma) in the Fayum Depression, Egypt.Akhnatenavus nefertiticyonsp. nov. has specialized, hypercarnivorous molars and an elongate cranial vault. InA. nefertiticyonthe tallest, piercing cusp on M1–M2is the paracone.Brychotherium ephalmosgen. et sp. nov. has more generalized molars that retain the metacone and complex talonids. InB. ephalmosthe tallest, piercing cusp on M1–M2is the metacone. We incorporate this new material into a series of phylogenetic analyses using a character-taxon matrix that includes novel dental, cranial, and postcranial characters, and samples extensively from the global record of the group. The phylogenetic analysis includes the first application of Bayesian methods to hyaenodont relationships.B. ephalmosis consistently placed within Teratodontinae, an Afro-Arabian clade with several generalist and hypercarnivorous forms, andAkhnatenavusis consistently recovered in Hyainailourinae as part of an Afro-Arabian radiation. The phylogenetic results suggest that hypercarnivory evolved independently three times within Hyaenodonta: in Teratodontinae, in Hyainailourinae, and in Hyaenodontinae. Teratodontines are consistently placed in a close relationship with Hyainailouridae (Hyainailourinae + Apterodontinae) to the exclusion of “proviverrines,” hyaenodontines, and several North American clades, and we propose that the superfamily Hyainailouroidea be used to describe this relationship. Using the topologies recovered from each phylogenetic method, we reconstructed the biogeographic history of Hyaenodonta using parsimony optimization (PO), likelihood optimization (LO), and Bayesian Binary Markov chain Monte Carlo (MCMC) to examine support for the Afro-Arabian origin of Hyaenodonta. Across all analyses, we found that Hyaenodonta most likely originated in Europe, rather than Afro-Arabia. The clade is estimated by tip-dating analysis to have undergone a rapid radiation in the Late Cretaceous and Paleocene; a radiation currently not documented by fossil evidence. During the Paleocene, lineages are reconstructed as dispersing to Asia, Afro-Arabia, and North America. The place of origin of Hyainailouroidea is likely Afro-Arabia according to the Bayesian topologies but it is ambiguous using parsimony. All topologies support the constituent clades–Hyainailourinae, Apterodontinae, and Teratodontinae–as Afro-Arabian and tip-dating estimates that each clade is established in Afro-Arabia by the middle Eocene.


2010 ◽  
Vol 84 (5) ◽  
pp. 935-954 ◽  
Author(s):  
Isaac S. Winkler ◽  
Conrad C. Labandeira ◽  
Torsten Wappler ◽  
Peter Wilf

Fossilized leaf mines and other traces of phytophagous insects provide a unique window into ecological and evolutionary associations of the past. Leaf-mining flies (Diptera: Agromyzidae) are an important component of the recent leaf-mining fauna, but their fossil record is sparse compared to other mining insect lineages; many putative agromyzid body fossils and traces are dubiously assigned. Agromyzid leaf mines often can be distinguished from those of other insects by the presence of an intermittent, fluidized frass trail that may alternate between the sides of the mine. Here, we describe two new Paleogene leaf mine fossils, Phytomyzites biliapchaensis Winkler, Labandeira and Wilf n. sp. from the early Paleocene of southeastern Montana, USA, occurring in leaves of Platanus raynoldsii (Platanaceae); and Phytomyzites schaarschmidti Wappler n. sp., from the middle Eocene of Messel, Germany, occurring in leaves of Toddalia ovata (Rutaceae). These fossils both exhibit frass trails indicative of an agromyzid origin, and P. biliapchaensis also exhibits associated stereotypical marks identical to damage caused by feeding punctures of extant adult female Agromyzidae prior to oviposition. Phytomyzites biliapchaensis represents the earliest confirmed record of Agromyzidae, and one of the earliest records for the large dipteran clade Schizophora. Plant hosts of both species belong to genera that are no longer hosts of leaf-mining Agromyzidae, suggesting a complex and dynamic history of early host-plant associations and, for the early Paleocene example, an evolutionary, possibly opportunistic colonization in the midst of the ecological chaos following the end-Cretaceous event in North America.


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