The South African stereospondyl Lydekkerina huxleyi (Tetrapoda, Temnospondyli) from the Lower Triassic of Australia

2006 ◽  
Vol 143 (6) ◽  
pp. 877-886 ◽  
Author(s):  
A. A. WARREN ◽  
R. DAMIANI ◽  
A. M. YATES
Keyword(s):  
South Africa ◽  
South African ◽  
Posterior Margin ◽  
Lower Triassic ◽  
Ventral Surface ◽  
The South ◽  
Karoo Basin ◽  
Skull Roof ◽  
Assemblage Zone

The first tetrapod fossil from the Rewan Formation of the Galilee Basin, central Queensland, Australia, is identified as Lydekkerina huxleyi, a stereospondyl found elsewhere only in the Lystrosaurus Assemblage Zone of South Africa. Apomorphies shared with L. huxleyi are: anterior palatal vacuity with anterodorsal projections from its posterior margin; ventral surface of skull roof with series of thickened ridges (condition unknown in other lydekkerinids); and vomerine shagreen present (possible autapomorphic reversal). Restudy of the only other Australian lydekkerinid, Chomatobatrachus halei, shows it to be distinct from L. huxleyi. The Rewan Formation, undifferentiated in the Galilee Basin, can be correlated with the Rewan Group of the Bowen Basin, and to the early part of the Lystrosaurus Assemblage Zone of the Karoo Basin, South Africa, which are of Griesbachian age. Varying palaeoenvironments may contribute to the contrasting nature of the Australian and South African faunas.

scholarly journals A new large cynodont from the Late Permian (Lopingian) of the South African Karoo Basin and its phylogenetic significance

2019 ◽  
Vol 186 (4) ◽  
pp. 983-1005 ◽  
Author(s):  
Fernando Abdala ◽  
Leandro C Gaetano ◽  
Roger M H Smith ◽  
Bruce S Rubidge
Keyword(s):  
South African ◽  
Posterior Margin ◽  
Cranial Morphology ◽  
Late Permian ◽  
Posterior Portion ◽  
Skull Length ◽  
Karoo Basin ◽  
Secondary Palate ◽  
Assemblage Zone

Abstract The Karoo Basin of South Africa has the best global record of Lopingian (Late Permian) non-mammaliaform cynodonts, currently represented by five species. We describe Vetusodon elikhulu gen. et sp. nov., documented by four specimens from the Daptocephalus Assemblage Zone. With a basal skull length of ~18 cm, it is the largest Lopingian cynodont and is also larger than Induan representatives of the group. Vetusodon elikhulu has a cranial morphology that departs notably from that previously documented for Permo-Triassic cynodonts. It features a short and extremely wide snout, resembling that of the contemporaneous therocephalian Moschorhinus, and has large incisors and canines that contrast with the small unicusped postcanines, suggesting a more important role of the anterior dentition for feeding. The dentary is extremely long and robust, with the posterior margin located closer to the craniomandibular joint than in other Lopingian and Induan cynodonts (e.g. Thrinaxodon). The secondary palate morphology of V. elikhulu is unique, being short and incomplete and with the posterior portion of the maxilla partly covering the vomer. A phylogenetic analysis suggests that V. elikhulu is the sister taxon of Eucynodontia and thus the most derived of the Lopingian to Induan cynodonts yet discovered.

Cranial morphology and phylogenetic relationships of the Middle Permian pareiasaur Embrithosaurus schwarzi from the Karoo Basin of South Africa

2019 ◽  
Author(s):  
Marc Johan Van den Brandt ◽  
Fernando Abdala ◽  
Bruce Sidney Rubidge
Keyword(s):  
South Africa ◽  
South African ◽  
Middle Permian ◽  
Cranial Morphology ◽  
Karoo Basin ◽  
Assemblage Zone ◽  
Mandibular Teeth ◽  
First Time ◽  
Globally Distributed ◽  
Distinct Taxon

Abstract Pareiasaurs were globally distributed, abundant, herbivorous parareptiles of the Middle to Late Permian, with the basal-most members found in the Middle Permian of South Africa. These basal taxa were particularly abundant and went extinct at the end of the Gaudalupian (Capitanian) at the top of the Tapinocephalus Assemblage Zone. Currently four taxa are recognized in this group: Bradysaurus seeleyi, B. baini, Nochelesaurus alexanderi and Embrithosaurus schwarzi, but they are all poorly understood. We here present the first detailed cranial description and updated diagnosis for Embrithosaurus schwarzi. No cranial autapomorphies were identified. However, Embrithosaurus schwarzi is a distinct taxon in this group, based on its unique dentition and using a combination of cranial features. It has nine marginal cusps on all maxillary and mandibular teeth, and wider maxillary teeth than in the co-occurring taxa, due to the marginal cusps being arranged more regularly around the crown, and the apex of the crown lacking the long, central, three-cusped trident. Our updated phylogenetic analysis recovers the four Middle Permian South African taxa as a monophyletic group for the first time, which we call Bradysauria, comprising a clade including Embrithosaurus, Bradysaurus baini and a polytomy including Nochelesaurus and Bradysaurus seeleyi.

SMALL FOOTPRINTS EXPAND MIDDLE PERMIAN AMPHIBIAN DIVERSITY IN THE SOUTH AFRICAN KAROO

Palaios ◽  
2020 ◽  
Vol 35 (1) ◽  
pp. 1-11 ◽  
Author(s):  
JUAN CARLOS CISNEROS ◽  
MICHAEL O. DAY ◽  
JACO GROENEWALD ◽  
BRUCE S. RUBIDGE
Keyword(s):  
South African ◽  
Middle Permian ◽  
Aquatic Biota ◽  
The South ◽  
Adult Form ◽  
Karoo Basin ◽  
Assemblage Zone

ABSTRACT Two new tetrapod trackways are described from the Tapinocephalus Assemblage Zone of the South African Karoo Basin. We interpret both to be traces attributable to small anamniote tetrapods. The larger footprints are tentatively referred to aff. Batrachichnus salamandroides. These imprints are distinguished from other records of Batrachichnus by a pentadactyl pes that produces only impressions of digits III–V. Digits I and II are recognized only by their drag marks. This trace occurs in association with a second set of footprints of uncertain affinities. However, these smaller imprints are not sufficiently well preserved and could represent undertracks or partially eroded footprints. None of the footprints can be attributed to the adult forms of the two temnospondyl taxa known from the Guadalupian part of the Karoo Basin: Rhinesuchus whaitsi or Rhinesuchoides tenuiceps. We interpret the aff. Batrachichnus trackway to have been produced by a small, adult temnospondyl or microsaur (Recumbirostrae), whereas the smaller set of footprints was likely made by a juvenile rhinesuchid or an unknown amphibian, either a paedomorphic form or a tiny adult form. The discovery shows that a more diverse aquatic biota existed at this time in the Karoo than osteological records currently suggest.

scholarly journals Cranial morphology of the middle Permian pareiasaur Nochelesaurus alexanderi from the Karoo Basin of South Africa

2021 ◽  
pp. 1-21
Author(s):  
Marc J. VAN DEN BRANDT ◽  
Bruce S. RUBIDGE ◽  
Julien BENOIT ◽  
Fernando ABDALA
Keyword(s):  
South Africa ◽  
South African ◽  
Local Group ◽  
Cranial Morphology ◽  
Karoo Basin ◽  
Lower Jaw ◽  
Basal Group ◽  
Assemblage Zone ◽  
Globally Distributed ◽  
Distinct Taxon

ABSTRACT Pareiasaurs were globally distributed, abundant, herbivorous parareptiles with the basal-most members found only in the mid-Permian of South Africa. These basal forms form a monophyletic group and were locally abundant and became extinct at the top of the Tapinocephalus Assemblage Zone at the end of the Guadalupian. Four species of basal pareiasaurs are currently recognised: Bradysaurus baini, B. seeleyi, Embrithosaurus schwarzi and Nochelesaurus alexanderi, but they are all poorly understood and there remains historic uncertainty as to their validity. In this paper, our second contribution designed to improve understanding of the basal group, we present the first detailed cranial description and updated diagnosis for Nochelesaurus alexanderi and demonstrate that it is a distinct taxon based on one cranial autapomorphy, a large transversely wide postparietal, and a combination of cranial characters. Within the local group of mid-Permian pareiasaurs, we recognise new dental features of Nochelesaurus alexanderi: non-symmetrical marginal cusp arrangements on upper and lower teeth resulting from an extra basal mesial cusp; an incipient horizontal cingulum on lower jaw teeth, sometimes with one or two tiny medial cingular cusps; and up to ten marginal cusps. Our study demonstrates that tooth morphology and orientation, cranial ornamentation, morphology of the cheek bosses, shape of the postfrontal and postparietal, and morphology of the distal paroccipital process of the opisthotic are the most useful to identify South African mid-Permian pareiasaurs.

scholarly journals The tetrapod fauna of the upper Permian Naobaogou Formation of China: 1. Shiguaignathus wangi gen. et sp. nov., the first akidnognathid therocephalian from China

PeerJ ◽  
2017 ◽  
Vol 5 ◽  
pp. e4150 ◽  
Author(s):  
Jun Liu ◽  
Fernando Abdala
Keyword(s):  
South Africa ◽  
Phylogenetic Analysis ◽  
South African ◽  
Fossil Record ◽  
Major Group ◽  
Upper Permian ◽  
The South ◽  
Karoo Basin ◽  
Basal Position

The Permian from China has a well-known terrestrial record where approximately 30 tetrapod taxa, including several therapsids, have been described. However, the record of therocephalians in China has remained elusive. Shiguaignathus wangi gen. et sp. nov., discovered in the Member III of the Naobaogou Formation, Nei Mongol, China, is here described. This is the first therocephalian recovered from this fauna and only the second from the Permian of China. It is represented by a well-preserved robust snout of a medium-sized animal. This is the first akidnognathid reported from the Chinese Permian and only the second genus from Laurasia as one genus is known from Russia whereas the remaining members of the group are from the South African Karoo Basin. A phylogenetic analysis of therocephalians supports a basal position of S. wangi within Akidnognathidae, followed by the Russian Annatherapsidus. Akidnognathidae is the latest major group of therocephalian appearing in the fossil record, and one of the few that does not have species from South Africa representing its most basal members.

Biostratigraphy of the Daptocephalus Assemblage Zone (Beaufort Group, Karoo Supergroup), South Africa

2020 ◽  
Vol 123 (2) ◽  
pp. 191-206 ◽  
Author(s):  
P.A. Viglietti
Keyword(s):  
South Africa ◽  
South African ◽  
Taxonomic Revision ◽  
Species Level ◽  
Karoo Basin ◽  
Assemblage Zone ◽  
Stratigraphic Ranges

Abstract The name Daptocephalus Assemblage Zone (DaAZ) is re-instated for vertebrate assemblages of the uppermost Permian strata (Balfour, upper Teekloof, and Normandien formations) of South Africa’s main Karoo Basin (MKB). This involved taxonomic revision of the dicynodontoid “Dicynodon” sensu lato, reviving Daptocephalus leoniceps, and revising the stratigraphic ranges of co-occurring index taxa (Theriognathus microps, Procynosuchus delaharpeae) of the Dicynodon Assemblage Zone (DiAZ) as it was known. This work has demonstrated the appearance of index taxa below the stratigraphically defined DiAZ. Moreover, the first appearance of Lystrosaurus maccaigi and Moschorhinus kitchingi in the upper reaches of the biozone calls for the establishment of a two-fold subdivision of the current DaAZ into lower (Dicynodon-Theriognathus) and upper (Lystrosaurus maccaigi-Moschorhinus) subzones. The biostratigraphic utility of Daptocephalus and other South African dicynodontoids outside of the MKB is limited due to basinal endemism at the species level and varying temporal ranges of dicynodontoids globally. Accordingly, their use is recommended only for correlation within the Karoo Basin at this stage.

First trematosaurid temnospondyl from the Lystrosaurus Assemblage Zone of South Africa and its biostratigraphic implications

2000 ◽  
Vol 137 (6) ◽  
pp. 659-665 ◽  
Author(s):  
ROSS DAMIANI ◽  
JOHANN NEVELING ◽  
JOHN HANCOX ◽  
BRUCE RUBIDGE
Keyword(s):  
South Africa ◽  
South African ◽  
Early Triassic ◽  
The South ◽  
Assemblage Zone

A large temnospondyl mandible from the Lystrosaurus Assemblage Zone (Early Triassic) of South Africa is referred to the higher-level taxon Trematosauridae. The mandible is remarkably similar to that described for Trematosaurus from the Middle Buntsandstein of Germany, a genus closely related to the South African Trematosuchus, for which the mandible is unknown. However, the mandible cannot be referred unequivocally to either of these taxa. Trematosuchus is considered to be restricted to the lowermost subzone of the overlying Cynognathus Assemblage Zone, which, based principally on the temnospondyl and therapsid fauna, is considered to be of Upper Olenekian age. The mandible described here extends back the range of the Trematosauridae to the underlying Lystrosaurus Assemblage Zone and argues against the long-held notion of a sharp palaeontological break between the faunas of the Lystrosaurus and Cynognathus Assemblage zones. It also supports the hypothesis that the Lystrosaurus Assemblage Zone extended up to the Upper Olenekian.

Adult morphology of the therocephalian Simorhinella baini from the middle Permian of South Africa and the taxonomy, paleobiogeography, and temporal distribution of the Lycosuchidae

2014 ◽  
Vol 88 (6) ◽  
pp. 1139-1153 ◽  
Author(s):  
Fernando Abdala ◽  
Christian F. Kammerer ◽  
Michael O. Day ◽  
Sifelani Jirah ◽  
Bruce S. Rubidge
Keyword(s):  
South Africa ◽  
South African ◽  
Type Species ◽  
Temporal Distribution ◽  
Ventral Surface ◽  
Middle Permian ◽  
Valid Species ◽  
Incertae Sedis ◽  
The Family ◽  
Assemblage Zone

The Middle Permian tetrapod fauna of the South African Beaufort Group is taxonomically diverse and includes representatives of all major therapsid groups, including the earliest records of Eutheriodontia. In the Middle Permian, eutheriodonts are represented mainly by large therocephalians, which made up a large proportion of the vertebrate predators in these faunas. Here we describe the skull and partial skeleton of a large therocephalian from the uppermost Tapinocephalus Assemblage Zone (AZ) of South Africa. A combination of features, including the short snout, presence of three to four upper postcanines and presence of teeth on the pterygoid processes, indicates that the new specimen belongs to the earliest-diverging therocephalian family, Lycosuchidae. The presence of a well-developed midline ridge on the ventral surface of the vomer indicates that the new specimen can be referred to Simorhinella baini, a species previously represented only by a tiny juvenile skull. The new specimen forms the basis for a taxonomic re-evaluation of the Lycosuchidae as well as of the geographic and stratigraphic range of the family. We recognize two valid species within the Lycosuchidae: the type species Lycosuchus vanderrieti represented by five specimens and Simorhinella baini represented by two specimens, with an additional 22 specimens currently identifiable as Lycosuchidae incertae sedis. Lycosuchid specimens range throughout the Tapinocephalus and Pristerognathus AZs; specimens of Simorhinella are restricted to the Tapinocephalus AZ, whereas Lycosuchus specimens are documented in both the Tapinocephalus and Pristerognathus AZs.

Foreigners' Rights to Work and Fair Labour Practices in South Africa: A Review of the South African Labour Legislation and Jurisprudence

2020 ◽  
Vol 35 (1) ◽  
Author(s):  
Andre Mangu
Keyword(s):  
South Africa ◽  
Human Rights ◽  
South African ◽  
Bill Of Rights ◽  
The South ◽  
Legislation And Jurisprudence ◽  
Foreign Nationals ◽  
Lay Persons ◽  
Democratic State ◽  
Labour Legislation

After several decades of apartheid rule, which denied human rights to the majority of the population on the ground of race and came to be regarded as a crime against humanity, South Africa adopted its first democratic Constitution in the early 1990s. The 1996 Constitution, which succeeded the 1993 interim Constitution, is considered one of the most progressive in the world. In its founding provisions, it states that South Africa is a democratic state founded on human dignity, the achievement of equality, the advancement of human rights and freedoms. The Constitution enshrines fundamental human rights in a justiciable Bill of Rights as a cornerstone of democracy. Unfortunately, in the eyes of a number of politicians, officials and lay-persons, the rights in the Bill of Rights accrue to South African citizens only. Xenophobia, which has been rampant since the end of apartheid, seems to support the idea that foreigners should not enjoy these rights. Foreign nationals have often been accused of posing a threat to South African citizens with regard to employment opportunities. In light of the South African legislation and jurisprudence, this article affirms the position of the South African labour law that foreign nationals are indeed protected by the Constitution and entitled to rights in the Bill of Rights, including the rights to work and fair labour practices.

The Engagement of Beneficiaries in Farm Governance of Restituted Land through the Communal Property Association Model: The Ideal Versus the Reality of Beneficiary Farms in South Africa

2019 ◽  
Vol 16 (2) ◽  
Author(s):  
Mokoko Piet Sebola ◽  
Malemela Angelinah Mamabolo
Keyword(s):  
South Africa ◽  
South African ◽  
Land Reform ◽  
Farm Management ◽  
The South ◽  
African Government ◽  
Communal Property ◽  
Farm Ownership ◽  
Land Restitution ◽  
Communal Property Associations

The purpose of this article is to evaluate the engagement of farm beneficiaries in South Africa in the governance of restituted farms through communal property associations. The South African government has already spent millions of rands on land restitution to correct the imbalance of the past with regard to farm ownership by the African communities. Various methods of farm management to benefit the African society have been proposed, however, with little recorded success. This article argues that the South African post-apartheid government was so overwhelmed by political victory in 1994 that they introduced ambitious land reform policies that were based on ideal thinking rather than on a pragmatic approach to the South African situation. We used qualitative research methods to argue that the engagement of farm beneficiaries in farm management and governance through communal property associations is failing dismally. We conclude that a revisit of the communal property associations model is required in order to strengthen the position of beneficiaries and promote access to land by African communities for future benefit.

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