scholarly journals Inbreeding parents should invest more resources in fewer offspring

2016 ◽  
Vol 283 (1843) ◽  
pp. 20161845 ◽  
Author(s):  
A. Bradley Duthie ◽  
Aline M. Lee ◽  
Jane M. Reid
Keyword(s):  
General Theory ◽  
Evolutionary Dynamics ◽  
Inbreeding Avoidance ◽  
The Other ◽  
Trade Off ◽  
Offspring Number ◽  
Focal Female ◽  
Special Cases ◽  
Selection For ◽  
Joint Consideration

Inbreeding increases parent–offspring relatedness and commonly reduces offspring viability, shaping selection on reproductive interactions involving relatives and associated parental investment (PI). Nevertheless, theories predicting selection for inbreeding versus inbreeding avoidance and selection for optimal PI have only been considered separately, precluding prediction of optimal PI and associated reproductive strategy given inbreeding. We unify inbreeding and PI theory, demonstrating that optimal PI increases when a female's inbreeding decreases the viability of her offspring. Inbreeding females should therefore produce fewer offspring due to the fundamental trade-off between offspring number and PI. Accordingly, selection for inbreeding versus inbreeding avoidance changes when females can adjust PI with the degree that they inbreed. By contrast, optimal PI does not depend on whether a focal female is herself inbred. However, inbreeding causes optimal PI to increase given strict monogamy and associated biparental investment compared with female-only investment. Our model implies that understanding evolutionary dynamics of inbreeding strategy, inbreeding depression, and PI requires joint consideration of the expression of each in relation to the other. Overall, we demonstrate that existing PI and inbreeding theories represent special cases of a more general theory, implying that intrinsic links between inbreeding and PI affect evolution of behaviour and intrafamilial conflict.

scholarly journals Inbreeding parents should invest more resources in fewer offspring

2016 ◽  
Author(s):  
A. Bradley Duthie ◽  
Aline M. Lee ◽  
Jane M. Reid
Keyword(s):  
General Theory ◽  
Evolutionary Dynamics ◽  
Inbreeding Avoidance ◽  
The Other ◽  
Trade Off ◽  
Offspring Number ◽  
Focal Female ◽  
Special Cases ◽  
Selection For ◽  
Joint Consideration

AbstractInbreeding increases parent-offspring relatedness and commonly reduces offspring viability, shaping selection on reproductive interactions involving relatives and associated parental investment (PI). Nevertheless, theories predicting selection for inbreeding versus inbreeding avoidance and selection for optimal PI have only been considered separately, precluding prediction of optimal PI and associated reproductive strategy given inbreeding. We unify inbreeding and PI theory, demonstrating that optimal PI increases when a female's inbreeding decreases the viability of her offspring. Inbreeding females should therefore produce fewer offspring due to the fundamental trade-off between offspring number and PI. Accordingly, selection for inbreeding versus inbreeding avoidance changes when females can adjust PI with the degree that they inbreed. In contrast, optimal PI does not depend on whether a focal female is herself inbred. However, inbreeding causes optimal PI to increase given strict monogamy and associated biparental investment compared to female-only investment. Our model implies that understanding evolutionary dynamics of inbreeding strategy, inbreeding depression, and PI requires joint consideration of the expression of each in relation to the other. Overall, we demonstrate that existing PI and inbreeding theories represent special cases of a more general theory, implying that intrinsic links between inbreeding and PI affect evolution of behaviour and intra-familial conflict.

scholarly journals Evolution of pre-copulatory and post-copulatory strategies of inbreeding avoidance and associated polyandry

2017 ◽  
Author(s):  
A. Bradley Duthie ◽  
Greta Bocedi ◽  
Ryan R. Germain ◽  
Jane M. Reid
Keyword(s):  
Inbreeding Depression ◽  
Evolutionary Dynamics ◽  
Lower Cost ◽  
Multiple Mating ◽  
Direct Costs ◽  
Inbreeding Avoidance ◽  
Evolutionary Constraints ◽  
Selection For ◽  
Evolution Of Polyandry ◽  
Or Modelling

AbstractInbreeding depression is widely hypothesised to drive adaptive evolution of pre-copulatory and post-copulatory mechanisms of inbreeding avoidance, which in turn are hypothesised to affect evolution of polyandry (i.e., female multiple mating). However, surprisingly little theory or modelling critically examines selection for pre-copulatory or post-copulatory inbreeding avoidance, or both strategies, given evolutionary constraints and direct costs, or examines how evolution of inbreeding avoidance strategies might feed back to affect evolution of polyandry. Selection for post-copulatory inbreeding avoidance, but not for pre-copulatory inbreeding avoidance, requires polyandry, while interactions between pre-copulatory and post-copulatory inbreeding avoidance might cause functional redundancy (i.e., ‘degeneracy’) potentially generating complex evolutionary dynamics among inbreeding strategies and polyandry. We used individual-based modelling to quantify evolution of interacting pre-copulatory and post-copulatory inbreeding avoidance and associated polyandry given strong inbreeding depression and different evolutionary constraints and direct costs. We found that evolution of post-copulatory inbreeding avoidance increased selection for initially rare polyandry, and that evolution of a costly inbreeding avoidance strategy became negligible over time given a lower cost alternative strategy. Further, fixed pre-copulatory inbreeding avoidance often completely precluded evolution of polyandry and hence post-copulatory inbreeding avoidance, but fixed post-copulatory inbreeding avoidance did not preclude evolution of pre-copulatory inbreeding avoidance. Evolution of inbreeding avoidance phenotypes and associated polyandry are therefore affected by evolutionary feedbacks and degeneracy. All else being equal, evolution of pre-copulatory inbreeding avoidance and resulting low polyandry is more likely when post-copulatory inbreeding avoidance is precluded or costly, and evolution of post-copulatory inbreeding avoidance greatly facilitates evolution of costly polyandry.

scholarly journals Black Hole Algorithm for Sustainable Design of Counterfort Retaining Walls

2020 ◽  
Vol 12 (7) ◽  
pp. 2767 ◽  
Author(s):  
Víctor Yepes ◽  
José V. Martí ◽  
José García
Keyword(s):  
Black Hole ◽  
Sustainable Design ◽  
Retaining Walls ◽  
The Other ◽  
Trade Off ◽  
Construction Company ◽  
Geometric Variables ◽  
The Stability ◽  
The Cost ◽  
Black Hole Algorithm

The optimization of the cost and CO 2 emissions in earth-retaining walls is of relevance, since these structures are often used in civil engineering. The optimization of costs is essential for the competitiveness of the construction company, and the optimization of emissions is relevant in the environmental impact of construction. To address the optimization, black hole metaheuristics were used, along with a discretization mechanism based on min–max normalization. The stability of the algorithm was evaluated with respect to the solutions obtained; the steel and concrete values obtained in both optimizations were analyzed. Additionally, the geometric variables of the structure were compared. Finally, the results obtained were compared with another algorithm that solved the problem. The results show that there is a trade-off between the use of steel and concrete. The solutions that minimize CO 2 emissions prefer the use of concrete instead of those that optimize the cost. On the other hand, when comparing the geometric variables, it is seen that most remain similar in both optimizations except for the distance between buttresses. When comparing with another algorithm, the results show a good performance in optimization using the black hole algorithm.

Security-Reliability Trade-Off Analysis of AN-Aided Relay Selection for Full-Duplex Relay Networks

2021 ◽  
Vol 70 (3) ◽  
pp. 2362-2377
Author(s):  
Zhanghua Cao ◽  
Xiaodong Ji ◽  
Jue Wang ◽  
Shibing Zhang ◽  
Yancheng Ji ◽  
...  
Keyword(s):  
Relay Selection ◽  
Relay Networks ◽  
Full Duplex ◽  
Trade Off ◽  
Selection For ◽  
Full Duplex Relay

scholarly journals Leader-chasing behavior in negative artificial triggered lightning flashes

2021 ◽  
Vol 11 (1) ◽  
Author(s):  
Fukun Wang ◽  
Jianguo Wang ◽  
Li Cai ◽  
Rui Su ◽  
Wenhan Ding ◽  
...  
Keyword(s):  
High Speed ◽  
The Other ◽  
Lightning Flash ◽  
High Speed Camera ◽  
Return Stroke ◽  
Catching Up ◽  
Special Cases ◽  
Triggered Lightning

AbstractTwo special cases of dart leader propagation were observed by the high-speed camera in the leader/return stroke sequences of a classical triggered lightning flash and an altitude-triggered lightning flash, respectively. Different from most of the subsequent return strokes preceded by only one leader, the return stroke in each case was preceded by two leaders occurring successively and competing in the same channel, which herein is named leader-chasing behavior. In one case, the polarity of the latter leader was opposite to that of the former leader and these two combined together to form a new leader, which shared the same polarity with the former leader. In the other case, the latter leader shared the same polarity with the former leader and disappeared after catching up with the former leader. The propagation of the former leader in this case seems not to be significantly influenced by the existence of the latter leader.

The effect of exposure to conspecifics on restlessness in the parasitoid wasp Nasonia vitripennis (Hymenoptera: Pteromalidae)

2007 ◽  
Vol 139 (5) ◽  
pp. 678-684 ◽  
Author(s):  
B.H. King
Keyword(s):  
Habitat Quality ◽  
Parasitoid Wasp ◽  
Poor Quality ◽  
The Other ◽  
Dispersal Ability ◽  
Nasonia Vitripennis ◽  
Adult Females ◽  
Selection For ◽  
The Difference ◽  
Abundance And Distribution

AbstractWhen habitat quality is variable, there should be strong selection for the ability to detect and respond to the variation. Adult females of the parasitoid wasp Nasonia vitripennis (Walker) are known to increase their restlessness (the proportion of time in locomotion) both during and after exposure to a poor quality host. Doing so provides a mechanism for leaving a poor host and potentially finding a better host. This study examined whether restlessness also changes in response to competition as indicated by the presence of adult conspecifics. Both restlessness and the probability of dispersing across an inhospitable environment were greater when a female was with another female than when she was alone. However, restlessness did not remain elevated after the other female was removed. In contrast with females, restlessness of males did not increase either during or after exposure to other males, and the probability of dispersing across an inhospitable environment was unaffected by the presence of another male. The difference between females and males may be related to differences in dispersal ability and in the abundance and distribution of hosts versus mates.

Identifying opportunities for value chain development in the Kenyan coffee sector

2018 ◽  
Vol 47 (2) ◽  
pp. 150-159 ◽  
Author(s):  
Emerta Aragie
Keyword(s):  
Value Chain ◽  
Export Performance ◽  
Export Subsidy ◽  
The Other ◽  
Cash Crop ◽  
Trade Off ◽  
Cash Crops ◽  
Standard Value ◽  
Coffee Sector ◽  
Productivity Gains

By developing a model that describes the Kenyan coffee value chain, this study evaluates opportunities emanating from four scenarios representing productivity gains in the various value chain stages of the coffee sector and additional three scenarios reflecting shifts in market situations. Results show that productivity-enhancing policies have stronger effects on coffee output and export performance if they target the milling stage of the value chain. Export subsidy and favourable external marketing conditions also have stronger effects, distributed comparably across the various value chain stages. We, however, found that these gains in the coffee sector come at the expense of other cash crops such as cotton, tea, sugar and tobacco. The approach followed in this study is relevant as this trade-off between coffee and the other cash crop sectors may not be visibly shown using standard value chain approaches.

The Circular Cylinder With a Band of Uniform Pressure on a Finite Length of the Surface

1941 ◽  
Vol 8 (3) ◽  
pp. A97-A104 ◽  
Author(s):  
M. V. Barton
Keyword(s):  
Circular Cylinder ◽  
Finite Length ◽  
Infinite Series ◽  
Fundamental Problem ◽  
The Other ◽  
Complete Picture ◽  
Uniform Pressure ◽  
Uniform Tension ◽  
Special Cases ◽  
Stress And Deformation

Abstract The solution to the fundamental problem of a cylinder with a uniform pressure over one half its length and a uniform tension on the other half is found by using the Papcovitch-Neuber solution to the general equations. In this paper, the results, given analytically in terms of infinite-series expressions, are exhibited as curves giving a complete picture of the stress and deformation. The case of a cylinder with a band of uniform pressure of any length, with the exception of very small ones, is then solved by the method of superposition. The stresses and displacements are evaluated for the special cases of a cylinder with a uniform pressure load of 1 diam and 1/2 diam in length. The problem of a cylinder heated over one half its length is solved by the same means.

Practical Aspects of Propulsion Shaft Alignment

2021 ◽  
Author(s):  
Kyriakos Avgouleas ◽  
Emmanouil Sarris ◽  
George Gougoulidis
Keyword(s):  
Cost Effectiveness ◽  
Numerical Model ◽  
High Speed ◽  
The Other ◽  
Technical Literature ◽  
Trade Off ◽  
Fea Model ◽  
Shaft Alignment ◽  
Alignment Strategy ◽  
Potential Damage

The economical and operational implications of poor alignment are indisputable for the propulsion shafting system of a commercial vessel. This holds true for naval vessels as well, although far less documented in the technical literature. This paper addresses some of the challenges associated with the proper alignment of a high-speed naval craft, which has been in service for many years. Laser bore-sighting was performed on a Guided Missile Fast Patrol Boat resting on a docking cradle. The measured bearing offsets were input to a FEA model of the shafting system to calculate bearing reactions and detect potential misalignment issues. Subsequent decisions regarding corrective measures take into account the results computed by the numerical model, experience from sister ships, the available documentation from the building yard and several other factors which are discussed in the paper. The solutions proposed are targeted towards a balanced trade-off between cost effectiveness and out-of-service time on one hand, and the risk of potential damage from misalignment on the other hand, which would seriously disrupt the ship’s operational availability. Practical aspects and lessons identified in the process are also presented, which demonstrate the distinct differences in alignment strategy of a high-speed naval craft compared to a typical commercial vessel.

COMPUTING THE RUPTURE DEGREE IN COMPOSITE GRAPHS

2010 ◽  
Vol 21 (03) ◽  
pp. 311-319 ◽  
Author(s):  
AYSUN AYTAC ◽  
ZEYNEP NIHAN ODABAS
Keyword(s):  
Complete Graph ◽  
Connected Graph ◽  
The Other ◽  
Trade Off ◽  
Number Of Components ◽  
Np Complete ◽  
Work Done ◽  
Better Than

The rupture degree of an incomplete connected graph G is defined by [Formula: see text] where w(G - S) is the number of components of G - S and m(G - S) is the order of a largest component of G - S. For the complete graph Kn, rupture degree is defined as 1 - n. This parameter can be used to measure the vulnerability of a graph. Rupture degree can reflect the vulnerability of graphs better than or independent of the other parameters. To some extent, it represents a trade-off between the amount of work done to damage the network and how badly the network is damaged. Computing the rupture degree of a graph is NP-complete. In this paper, we give formulas for the rupture degree of composition of some special graphs and we consider the relationships between the rupture degree and other vulnerability parameters.

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