scholarly journals Coarse woody debris in the old-growth forests of British Columbia

2003 ◽  
Vol 11 (S1) ◽  
pp. S135-S157 ◽  
Author(s):  
M C Feller
Keyword(s):  
British Columbia ◽  
Coarse Woody Debris ◽  
Functional Role ◽  
Animal Species ◽  
Woody Debris ◽  
Douglas Fir ◽  
Forest Age ◽  
Old Growth ◽  
Old Growth Forests

This paper synthesizes data extracted from the literature and data collected in various studies by the author on the quantity, characteristics, and functional importance of coarse woody debris (CWD) in the old-growth forests of British Columbia (B.C.). There is little agreement in the literature about the minimum diameter of CWD or the number of decay classes recognized. In western North America, five decay classes are commonly used, but recent studies suggest fewer decay classes are preferable. Comparisons among decay classes and biogeoclimatic zones and subzones in B.C. reveal that quantities and volumes are greatest (up to approximately 60 kg/m2 and approximately 1800 m3/ha, respectively), and CWD persists the longest (sometimes in excess of 1000 years) in the Coastal Western Hemlock (CWH) biogeoclimatic zone. The quantity and ground cover of CWD increase with forest productivity. Persistence of CWD has varied from less than 100 to over 800 years in two coastal (CWH and Mountain Hemlock (MH)) and three interior (Interior Douglas-fir (IDF), Interior Cedar–Hemlock (ICH), and Engelmann Spruce – Subalpine Fir (ESSF)) biogeoclimatic zones. Trends in CWD quantity with forest age in managed coastal B.C. forests suggest a U-shaped curve, with greater quantities occurring in recent cutovers than in old-growth forests, and lowest quantities occurring in middle-aged forests. This may be the normal trend in CWD with forest age, with departures from this trend resulting from disturbance- or environment-specific factors. Relatively large amounts of data exist on the characteristics of CWD in the CWH, IDF, ICH, ESSF, and Boreal White and Black Spruce (BWBS) biogeoclimatic zones, but such data for the Coastal Douglas-fir, Sub-Boreal Pine–Spruce, Sub-Boreal Spruce (SBS), and Spruce–Willow–Birch biogeoclimatic zones appear relatively sparse. There have been few studies of the functional role of CWD in B.C. forests, but those studies that have been completed indicate that CWD is an important habitat component for some plant and animal species. A total of 169 plant species, including >95% of all lichens and liverworts, were found to grow on CWD in old-growth forests in the CWH, MH, IDF, ICH, and ESSF biogeoclimatic zones. One third of these species were restricted to CWD. Studies in several biogeoclimatic zones have found that CWD provided preferred habitat for and was associated with higher populations of some small animal species, such as shrews, some voles, and some salamanders, in old-growth forests, but the effects varied with species and biogeoclimatic zone. The nutrient cycling role of CWD is not yet well known, but it currently appears to be relatively insignificant in B.C. old-growth forests. Although it has been considered that CWD could increase mineral soil acidification and eluviation, no evidence for this was found in a study of the CWH, MH, IDF, ICH, ESSF, BWBS, and SBS biogeoclimatic zones. Future studies of the functional role of CWD should consider both scale (square metre vs. hectare) and temporal (changes in CWD with forest age) issues, as studies including these are sparse and both may be important. Key words: biogeoclimatic zones, British Columbia, coarse woody debris, old-growth forests.

scholarly journals Bryophyte Conservation in Managed Boreal Landscapes: Fourteen-Year Impacts of Partial Cuts on Epixylic Bryophytes

2021 ◽  
Vol 4 ◽  
Author(s):  
Jeffrey Opoku-Nyame ◽  
Alain Leduc ◽  
Nicole J. Fenton
Keyword(s):  
Species Composition ◽  
Coarse Woody Debris ◽  
Black Spruce ◽  
Woody Debris ◽  
Permanent Plots ◽  
Old Growth ◽  
Clear Cut ◽  
Old Growth Forest ◽  
Old Growth Forests ◽  
Over Time

Clear cut harvest simplifies and eliminates old growth forest structure, negatively impacting biodiversity. Partial cut harvest has been hypothesized (1) to have less impact on biodiversity than clear cut harvest, and (2) to encourage old growth forest structures. Long-term studies are required to test this hypothesis as most studies are conducted soon after harvest. Using epixylic bryophytes as indicators, this study addresses this knowledge gap. Fourteen years after harvest, we examined changes in epixylic bryophyte community composition richness and traits, and their microhabitats (coarse woody debris characteristics and microclimate) along an unharvested, partial cuts and clear cuts harvest treatment in 30 permanent plots established in the boreal black spruce (Picea mariana) forests of northwestern Quebec, Canada. Our results were compared to those of an initial post-harvest study (year 5) and to a chronosequence of old growth forests to examine species changes over time and the similarity of bryophyte communities in partial cut and old growth forests. Coarse woody debris (CWD) volume by decay class varied among harvest treatments with partial cuts and clear cuts recording lower volumes of early decay CWD. The epixylic community was richer in partial cuts than in mature unharvested forests and clear cuts. In addition, species richness and overall abundance doubled in partial and clear cuts between years 5 and 14. Species composition also differed among treatments between years 5 and 14. Furthermore, conditions in partial cut stands supported small, drought sensitive, and old growth confined species that are threatened by conditions in clear cut stands. Lastly, over time, species composition in partial cuts became more similar to old growth forests. Partial cuts reduced harvest impacts by continuing to provide favorable microhabitat conditions that support epixylic bryophytes. Also, partial cut harvest has the potential to encourage old growth species assemblages, which has been a major concern for biodiversity conservation in managed forest landscapes. Our findings support the promotion of partial cut harvest as an effective strategy to achieve species and habitat conservation goals.

Occurrence of Piloderma fallax in young, rotation-age, and old-growth stands of Douglas-fir (Pseudotsuga menziesii) in the Cascade Range of Oregon, U.S.A.

2000 ◽  
Vol 78 (8) ◽  
pp. 995-1001 ◽  
Author(s):  
J E Smith ◽  
R Molina ◽  
M MP Huso ◽  
M J Larsen
Keyword(s):  
Forest Management ◽  
Pseudotsuga Menziesii ◽  
Ectomycorrhizal Fungi ◽  
Coarse Woody Debris ◽  
Woody Debris ◽  
Douglas Fir ◽  
Percent Cover ◽  
Old Growth ◽  
Decay Class ◽  
Rotation Age

Yellow mycelia and cords of Piloderma fallax (Lib.) Stalp. were more frequently observed in old-growth stands than in younger managed stands of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco). Piloderma fallax frequency and percent cover data were collected from 900 plots in three replicate stands in each of three forest age classes over 2 years in both spring and fall. Piloderma fallax is strongly associated with stand age; it occurred in 57% of plots in old-growth, 6% of rotation-age, and 1% of young stands. Presence of Piloderma fallax was related to the percent cover of coarse woody debris (CWD) in decay class 5. Piloderma fallax was approximately 2.5 times more likely to occur in a plot with CWD decay class 5 present than in plots without. The probability that it would occur in a plot increased by approximately 20% for every 10% increase in percent cover of CWD decay class 5. However, the percent cover of Piloderma fallax was not strongly related to the percent cover of CWD in decay class 5. Frequency of occurrence did not differ among sampling times. Occurrence of Piloderma fallax may indicate suitable substrate for ectomycorrhizal fungi associated with CWD and may be important in forest management for the maintenance of biodiversity and old-growth components in young managed stands.Key words: Piloderma fallax, coarse woody debris, Pseudotsuga menziesii, forest management, ectomycorrhizal fungi, biodiversity.

Coarse woody debris and soil respiration in a clearcut on the Olympic Peninsula, Washington, U.S.A.

1996 ◽  
Vol 26 (8) ◽  
pp. 1337-1345 ◽  
Author(s):  
James L. Marra ◽  
Robert L. Edmonds
Keyword(s):  
Soil Respiration ◽  
Respiration Rate ◽  
Coarse Woody Debris ◽  
Woody Debris ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Olympic Peninsula ◽  
Old Growth ◽  
Growth Site ◽  
Respiration Rates

Coarse woody debris (CWD) and soil respiration rates were measured using soda lime traps on a clearcut site in the Hoh River Valley on the west side of the Olympic Peninsula, Washington. The influence of species of CWD (western hemlock (Tsugaheterophylla (Raf.) Sarg.) and Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco)), decay class, and log diameter on respiration rates was determined. CWD and soil respiration were measured every 4 weeks from October 1991 to November 1992 along with CWD and soil temperature and moisture contents. Western hemlock logs respired at a significantly higher rate (4.05 g CO2•m−2•day−1) than Douglas-fir logs (2.94 g CO2•m−2•day−1). There were no significant differences between respiration rates for decay classes 1–2, 3, and 5 logs (4.47, 3.69, and 4.28 g CO2•m−2•day−1, respectively), and there was no strong relationship between CWD respiration rate and log diameter. The highest average respiration rate was from the soil in the clearcut (5.22 g CO2•m−2•day−1). Averaged for the year, log and soil respiration rates in the clearcut site were similar to those in an adjacent old-growth forested site. However, seasonal fluctuations were greater on the clearcut site. Higher summer respiration rates and lower winter rates observed on the clearcut relative to the old-growth site appeared to be driven more by temperature than by moisture. Clear-cutting also resulted in higher summer CWD and soil temperatures and lower winter temperatures compared with the old-growth site.

Comparison of structural and compositional characteristics in old-growth and mature, managed hardwood forests of Minnesota, U.S.A.

1999 ◽  
Vol 29 (10) ◽  
pp. 1479-1489 ◽  
Author(s):  
C M Hale ◽  
J Pastor ◽  
K A Rusterholz
Keyword(s):  
Species Composition ◽  
Plant Species ◽  
Coarse Woody Debris ◽  
Woody Debris ◽  
Structural Features ◽  
Hardwood Forests ◽  
Plant Species Composition ◽  
Managed Forests ◽  
Old Growth ◽  
Old Growth Forests

Extended rotation of managed temperate hardwood forests is sometimes presumed to provide the important compositional and structural features of old-growth hardwood forests. However, the features of temperate hardwood old-growth and managed stands of extended rotation age have not been fully quantified and compared. This study compared quantitative parameters (density and volume of logs and snags, coarse woody debris volume (volume of logs + volume of snags), the proportion of hollow logs, basal area and tree, sapling, large seedling and small seedling densities), distributional patterns (diameter class and rot class of live trees, decay class of logs), and vascular plant species composition and diversity in old-growth and mature, managed sugar maple (Acer saccharum L.) - basswood (Tilia americana L.) and northern red oak (Quercus rubra L.) stands. Old-growth forests had higher coarse woody debris volumes and higher proportions of hollow logs, of live trees in large diameter classes, of logs in decay classes 1 and 2, and of live trees in rot classes 3-5 than the mature, managed forests. Old-growth and mature, managed forests did not differ significantly in plant species composition. These results indicate that, while older extended-rotation, managed stands can be very similar compositionally to old-growth forests, they differ quantitatively in structural features.

scholarly journals Maintaining attributes of old-growth forests in coastal B.C. through variable retention

2003 ◽  
Vol 79 (3) ◽  
pp. 570-578 ◽  
Author(s):  
W J Beese ◽  
B G Dunsworth ◽  
K. Zielke ◽  
B. Bancroft
Keyword(s):  
British Columbia ◽  
Pacific Northwest ◽  
Biological Diversity ◽  
Woody Debris ◽  
Structural Complexity ◽  
Structural Features ◽  
Management Program ◽  
Old Growth ◽  
Variable Retention ◽  
Old Growth Forests

Variable retention is a new approach to harvesting and silvicultural systems that was developed by ecologists in the Pacific Northwest region of North America to address a wide array of forest management goals. Variable retention recognizes that natural disturbances, such as fire, wind or disease, nearly always leave some standing structure from the original forest. This structural complexity plays an important role in forest ecosystem function and biological diversity. A new "retention silvicultural system" was defined that leaves trees distributed throughout harvested areas. This system facilitates retention of structural features of old-growth forests, such as live and dead trees of varying sizes, multiple canopy layers, and coarse woody debris. Weyerhaeuser's British Columbia Coastal Group will use the variable retention approach for all harvesting by 2003. More than 75% of the company's coastal harvesting in British Columbia used variable retention in 2001. Company guidelines describe the amount, type, and spatial distribution of retention for groups and individual trees. An adaptive management program is monitoring the amount and type of structural attributes retained in relation to the original forest. Key words: old-growth forests, variable retention, silvicultural systems, biodiversity, landscape zoning

The role of coarse woody debris in forest aquatic habitats: Implications for management

1995 ◽  
Vol 5 (2) ◽  
pp. 143-166 ◽  
Author(s):  
A. M. Gurnell ◽  
K. J. Gregory ◽  
G. E. Petts
Keyword(s):  
Coarse Woody Debris ◽  
Woody Debris ◽  
Aquatic Habitats

Coarse Woody Debris in Douglas-Fir Forests of Western Oregon and Washington

Ecology ◽  
1988 ◽  
Vol 69 (6) ◽  
pp. 1689-1702 ◽  
Author(s):  
Thomas A. Spies ◽  
Jerry F. Franklin ◽  
Ted B. Thomas
Keyword(s):  
Coarse Woody Debris ◽  
Woody Debris ◽  
Douglas Fir

Astrosclereids In Needles Of Old-Growth Douglas-Fir Trees

1999 ◽  
Vol 5 (S2) ◽  
pp. 1240-1241
Author(s):  
M. Apple ◽  
A. Soeldner ◽  
R. Hamill ◽  
K. Tiekotfer
Keyword(s):  
Pacific Northwest ◽  
Cell Walls ◽  
Douglas Fir ◽  
Old Growth ◽  
Mesophyll Cells ◽  
Mesophyll Tissue ◽  
The Pacific ◽  
And Function ◽  
Development Structure

Old-growth Douglas-fir trees in the Pacific Northwest are venerable giants that often live for 500 years and reach heights of over 75 meters. Their needles are relatively ephemeral and small but have the important role of interacting with the atmosphere in order to transpire and photosynthesize. Within the photosynthetic mesophyll tissue of Douglas-fir needles, there are large, non-living cells with lignified secondary cell walls that are known as astrosclereids. Apparent channels in the secondary wall may provide a route for exchange or transport of materials between the astrosclereid lumen and mesophyll cells or the vascular cylinder. Astrosclereids may be involvev d in storage of secondary metabolites such as tannin and may develop in response to fungi, mistletoe, or other pathogens. More knowledge is needed about the development, structure and function of astrosclereids.Needles were collected from sapling and old-growth Douglas-fir, Pseudotsuga menziesii, (Mirb.) Franco, trees at the Wind River Canopy Crane in Carson, Washington and from three sites in the Cascade Mountains of Oregon in 1997 and 1998.

Coarse woody debris in old-growth temperate beech (Nothofagus) forests of New Zealand

1994 ◽  
Vol 24 (10) ◽  
pp. 1989-1996 ◽  
Author(s):  
Glenn H. Stewart ◽  
Larry E. Burrows
Keyword(s):  
New Zealand ◽  
Coarse Woody Debris ◽  
Tree Mortality ◽  
Structural Integrity ◽  
Woody Debris ◽  
Hardwood Forests ◽  
Old Growth ◽  
Carbon And Nitrogen ◽  
Decay Sequence ◽  
Nothofagus Forests

The volume, biomass, and carbon and nitrogen content of coarse woody debris were measured on three 1-ha reference plots in old-growth Nothofagusfusca (Hook. f.) Oerst.–Nothofagusmenziesii (Hook. f.) Oerst. forest on the South Island of New Zealand. Two decay sequences for logs and one for standing dead trees (snags) were recognised from two-way indicator species analysis (TWINSPAN) of up to 30 variables related to physical characteristics and structural integrity. Wood volume (up to 800 m3•ha−1) and biomass were high (up to 300 Mg•ha−1), and the inside-out decay sequence from heartwood to sapwood was unusual compared with that of other temperate hardwood forests. Coarse woody debris represented significant carbon and nitrogen pools, with ca. 150 Mg•ha−1 and 370 kg•ha−1, respectively, in one stand. The coarse woody debris component of these broad-leaved evergreen hardwood forests was much higher than that reported for other temperate hardwood forests and approaches that of many northern hemisphere conifer forests. The large coarse woody debris pools are discussed in relation to live stand biomass, natural disturbances and tree mortality, and decomposition processes.

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