Revision of Dobodura Darlington (Coleoptera: Carabidae: Odacanthini): Diversification on accreted terranes of northern New Guinea

The Papuan endemic genus Dobodura Darlington is taxonomically revised, with five newly described species — Dobodura alildablldooya sp. n., D. hexaspina sp. n., D. obtusa sp. n., D. svensoni sp. n., and D. toxopei sp. n. — complementing the type species, D. armata Darlington. The sympatric Dobodura alildablldooya and D. svensoni are described from Chimbu Province, Papua New Guinea. Known distributions of the other three new species are: D. hexaspina, Madang Province, P.N.G.; D. obtusa, Olsobip, Fly River, Western Province, P.N.G.; and D. toxopei, Bernhard Camp, Papua, Indonesia. Dobodura is the sole precinctive Papuan genus in an Australian-Papuan clade also including Clarencia Sloane, Dicraspeda Chaudoir, and Eudalia Laporte. Phylogenetic analysis of Dobodura places its known earliest divergence event on the northern New Guinea margin of the Australian craton. Later divergence events result in species occupying island-arc terranes progressively incorporated into present-day northern New Guinea, commencing in the Miocene.