How rare is phyletic gradualism and what is its evolutionary significance? Evidence from Jurassic bivalves

Paleobiology ◽  
1978 ◽  
Vol 4 (1) ◽  
pp. 16-25 ◽  
Author(s):  
Anthony Hallam
Keyword(s):  
Sea Level ◽  
High Stress ◽  
Adaptive Strategy ◽  
Dominant Mode ◽  
Species Selection ◽  
Size Increase ◽  
Evolutionary Significance ◽  
Punctuated Equilibria ◽  
Sea Levels ◽  
Speciation Rates

Both intensive analysis of Gryphaea and an extensive survey of Jurassic bivalves in Europe supports the punctuated equilibria model of speciation, with the exception of phyletic size increase, which appears within the limits of the data to be gradualistic and affects at least a significant minority of species. The size increase takes place both within species and between successive species in a lineage. In Liassic Gryphaea, a combination of allometry in the ontogenetic development of the ancestral species and paedomorphosis led also to changes of shape up the sequence. The production of evolutionary trends by species selection is not supported.The favoured interpretation of phyletic size increase is a gradual delay in maturation time consequent upon a change in the organisms' adaptive strategy from the r-selected to the K-selected mode and is backed up by evidence on changing population numbers and distribution. An extinction/speciation model is proposed for neritic organisms based on fluctuating sea levels. Times of low sea level or regression correspond with times of high stress and hence high extinction among stenotopic organisms and increased rates of allopatric speciation, with r-selection as the dominant mode. Times of high sea level or transgression correspond with low extinction and speciation rates and increased freedom of migration. K-selection is the dominant mode and often leads to phyletic size increase provided the environment remains stable for a sufficiently long period.

Patterns of speciation in Jurassic Gryphaea

Paleobiology ◽  
1982 ◽  
Vol 8 (4) ◽  
pp. 354-366 ◽  
Author(s):  
Anthony Hallam
Keyword(s):  
Growth Rate ◽  
Physical Environment ◽  
World Wide ◽  
Species Selection ◽  
Size Increase ◽  
Initial Increase ◽  
Bivalve Species ◽  
Punctuated Equilibria ◽  
And Migration

It is shown that the abundant Jurassic oyster Gryphaea provides material adequate to test biogeographic and evolutionary hypotheses involving speciation. Phyletic size increase is widespread if not characteristic and tends to be marked by a sharp initial increase followed by a longer period of stability. The larger size of phyletically younger forms is apparently because of greater age of individuals rather than a higher growth rate. Species durations fall within the range of the majority of Jurassic bivalve species. The detailed study of European Gryphaea indicates a pattern of punctuated equilibria allied with morphological trends, some of which are paedomorphic. Gradualistic, species selection and cladistic models of change are not supported. World-wide analysis of Gryphaea species supports centres of origin and migration rather than vicariance, and a relationship exists between migrational and evolutionary events and changes in the physical environment.

Punctuated equilibria: the tempo and mode of evolution reconsidered

Paleobiology ◽  
1977 ◽  
Vol 3 (2) ◽  
pp. 115-151 ◽  
Author(s):  
Stephen Jay Gould ◽  
Niles Eldredge
Keyword(s):  
Fossil Record ◽  
A Priori ◽  
Dominant Mode ◽  
Species Selection ◽  
Geological History ◽  
Genetic Changes ◽  
Punctuated Equilibria ◽  
Apparent Direction ◽  
History Of ◽  
Fair Assessment

We believe that punctuational change dominates the history of life: evolution is concentrated in very rapid events of speciation (geologically instantaneous, even if tolerably continuous in ecological time). Most species, during their geological history, either do not change in any appreciable way, or else they fluctuate mildly in morphology, with no apparent direction. Phyletic gradualism is very rare and too slow, in any case, to produce the major events of evolution. Evolutionary trends are not the product of slow, directional transformation within lineages; they represent the differential success of certain species within a clade—speciation may be random with respect to the direction of a trend (Wright's rule).As an a priori bias, phyletic gradualism has precluded any fair assessment of evolutionary tempos and modes. It could not be refuted by empirical catalogues constructed in its light because it excluded contrary information as the artificial result of an imperfect fossil record. With the model of punctuated equilibria, an unbiased distribution of evolutionary tempos can be established by treating stasis as data and by recording the pattern of change for all species in an assemblage. This distribution of tempos can lead to strong inferences about modes. If, as we predict, the punctuational tempo is prevalent, then speciation—not phyletic evolution—must be the dominant mode of evolution.We argue that virtually none of the examples brought forward to refute our model can stand as support for phyletic gradualism; many are so weak and ambiguous that they only reflect the persistent bias for gradualism still deeply embedded in paleontological thought. Of the few stronger cases, we concentrate on Gingerich's data for Hyopsodus and argue that it provides an excellent example of species selection under our model. We then review the data of several studies that have supported our model since we published it five years ago. The record of human evolution seems to provide a particularly good example: no gradualism has been detected within any hominid taxon, and many are long-ranging; the trend to larger brains arises from differential success of essentially static taxa. The data of molecular genetics support our assumption that large genetic changes often accompany the process of speciation.Phyletic gradualism was an a priori assertion from the start—it was never “seen” in the rocks; it expressed the cultural and political biases of 19th century liberalism. Huxley advised Darwin to eschew it as an “unnecessary difficulty.” We think that it has now become an empirical fallacy. A punctuational view of change may have wide validity at all levels of evolutionary processes. At the very least, it deserves consideration as an alternate way of interpreting the history of life.

scholarly journals Economic damages from Hurricane Sandy attributable to sea level rise caused by anthropogenic climate change

2021 ◽  
Vol 12 (1) ◽  
Author(s):  
Benjamin H. Strauss ◽  
Philip M. Orton ◽  
Klaus Bittermann ◽  
Maya K. Buchanan ◽  
Daniel M. Gilford ◽  
...  
Keyword(s):  
Climate Change ◽  
Sea Level Rise ◽  
Sea Level ◽  
East Coast ◽  
The United States ◽  
Hurricane Sandy ◽  
Water Levels ◽  
Anthropogenic Climate Change ◽  
Economic Damage ◽  
Sea Levels

AbstractIn 2012, Hurricane Sandy hit the East Coast of the United States, creating widespread coastal flooding and over $60 billion in reported economic damage. The potential influence of climate change on the storm itself has been debated, but sea level rise driven by anthropogenic climate change more clearly contributed to damages. To quantify this effect, here we simulate water levels and damage both as they occurred and as they would have occurred across a range of lower sea levels corresponding to different estimates of attributable sea level rise. We find that approximately $8.1B ($4.7B–$14.0B, 5th–95th percentiles) of Sandy’s damages are attributable to climate-mediated anthropogenic sea level rise, as is extension of the flood area to affect 71 (40–131) thousand additional people. The same general approach demonstrated here may be applied to impact assessments for other past and future coastal storms.

Records of sea-level highstand over the Meghalayan age/late Holocene from uranium-series ages of beachrock in Weizhou Island, northern South China Sea

The Holocene ◽  
2021 ◽  
pp. 095968362110332
Author(s):  
Tingli Yan ◽  
Kefu Yu ◽  
Rui Wang ◽  
Wenhui Liu ◽  
Leilei Jiang
Keyword(s):  
South China Sea ◽  
Sea Level ◽  
South China ◽  
Intertidal Zone ◽  
Meteoric Water ◽  
Northern South China Sea ◽  
Water Environment ◽  
Sea Levels ◽  
China Sea ◽  
Micro Structures

Beachrock is considered a good archive for past sea-levels because of its unique formation position (intertidal zone). To evaluate sea-level history in the northern South China Sea, three well-preserved beachrock outcrops (Beigang, Gongshanbei, and Hengling) at Weizhou Island, northern South China Sea were selected to examine their relative elevation, sedimentological, mineralogical, and geochemical characteristics. Acropora branches with well-preserved surface micro-structures were selected from the beachrocks and used to determine the ages of these beachrocks via U-series dating. The results show that the beachrocks are composed of coral reef sediments, terrigenous clastics, volcanic clastics, and various calcite cements. These sediments accumulated in the intertidal zone of Weizhou Island were then cemented in a meteoric water environment. The U-series ages of beachrocks from Beigang, Gongshanbei, and Hengling are 1712–768 ca. BP, 1766–1070 ca. BP, and 1493–604 ca. BP (before 1950 AD) respectively. Their elevations are 0.91–1.16 m, 0.95–1.24 m, and 0.82–1.17 m higher than the modern homologous sedimentary zones, respectively. Therefore, we concluded that the sea-level in the Meghalayan age (1766–604 ca. BP) was 0.82–1.24 m higher than the present, and that the sea-level over this period showed a declining trend.

scholarly journals Flooding Conditions at Aveiro Port (Portugal) within the Framework of Projected Climate Change

2021 ◽  
Vol 9 (6) ◽  
pp. 595
Author(s):  
Américo Soares Ribeiro ◽  
Carina Lurdes Lopes ◽  
Magda Catarina Sousa ◽  
Moncho Gomez-Gesteira ◽  
João Miguel Dias
Keyword(s):  
Climate Change ◽  
Sea Level ◽  
Climate Change Impacts ◽  
Storm Surges ◽  
Historical Period ◽  
Return Periods ◽  
Wave Regime ◽  
Mean Sea Level ◽  
Sea Levels ◽  
Far Future

Ports constitute a significant influence in the economic activity in coastal areas through operations and infrastructures to facilitate land and maritime transport of cargo. Ports are located in a multi-dimensional environment facing ocean and river hazards. Higher warming scenarios indicate Europe’s ports will be exposed to higher risk due to the increase in extreme sea levels (ESL), a combination of the mean sea level, tide, and storm surge. Located on the west Iberia Peninsula, the Aveiro Port is located in a coastal lagoon exposed to ocean and river flows, contributing to higher flood risk. This study aims to assess the flood extent for Aveiro Port for historical (1979–2005), near future (2026–2045), and far future (2081–2099) periods scenarios considering different return periods (10, 25, and 100-year) for the flood drivers, through numerical simulations of the ESL, wave regime, and riverine flows simultaneously. Spatial maps considering the flood extent and calculated area show that most of the port infrastructures' resilience to flooding is found under the historical period, with some marginal floods. Under climate change impacts, the port flood extent gradually increases for higher return periods, where most of the terminals are at high risk of being flooded for the far-future period, whose contribution is primarily due to mean sea-level rise and storm surges.

scholarly journals Twenty-first century response of Petermann Glacier, northwest Greenland to ice shelf loss

2020 ◽  
pp. 1-11
Author(s):  
Emily A. Hill ◽  
G. Hilmar Gudmundsson ◽  
J. Rachel Carr ◽  
Chris R. Stokes ◽  
Helen M. King
Keyword(s):  
Sea Level ◽  
First Century ◽  
Ice Shelf ◽  
Ice Shelves ◽  
Sea Levels ◽  
Grounding Line ◽  
Global Mean Sea Level ◽  
Near Future ◽  
Global Mean ◽  
Large Sections

Abstract Ice shelves restrain flow from the Greenland and Antarctic ice sheets. Climate-ocean warming could force thinning or collapse of floating ice shelves and subsequently accelerate flow, increase ice discharge and raise global mean sea levels. Petermann Glacier (PG), northwest Greenland, recently lost large sections of its ice shelf, but its response to total ice shelf loss in the future remains uncertain. Here, we use the ice flow model Úa to assess the sensitivity of PG to changes in ice shelf extent, and to estimate the resultant loss of grounded ice and contribution to sea level rise. Our results have shown that under several scenarios of ice shelf thinning and retreat, removal of the shelf will not contribute substantially to global mean sea level (<1 mm). We hypothesize that grounded ice loss was limited by the stabilization of the grounding line at a topographic high ~12 km inland of its current grounding line position. Further inland, the likelihood of a narrow fjord that slopes seawards suggests that PG is likely to remain insensitive to terminus changes in the near future.

Mean Sea Level as a Reference for Geodetic Leveling

1974 ◽  
Vol 28 (5) ◽  
pp. 524-530 ◽  
Author(s):  
G. W. Lennon
Keyword(s):  
Sea Level ◽  
World Ocean ◽  
Equipotential Surface ◽  
Scientific Discipline ◽  
Mean Sea Level ◽  
Sea Levels ◽  
Coastal Sea ◽  
Long Time ◽  
Marine System

The use of mean sea level as a surface of reference that might provide an independent control for geodetic leveling has been a long term goal arising from the classical analogy between the geoid as an equipotential surface and the surface assumed by a hypothetical undisturbed world ocean. The problems associated with this aim are now known to be vast, and are associated with the dynamics of the marine system, notably its response to meteorological forces, to variations in density and to the effects of basic circulation patterns. In consequence the mean sea level surface varies rapidly in both time and space. This identifies in fact a distinctive scientific discipline, coastal geodesy, in which contributions are required by both geodesists and oceanographers. It has come to be recognized that the coastal zone is a hazardous environment for all observational techniques concerned. On the one hand, the difficulties of measurement of coastal sea levels have only recently been understood; on the other hand, precise leveling procedures are now known to be influenced by the attraction of marine tides and by crustal deformation of tidal loading. Much of the data available for study are therefore inadequate and, moreover, it should be noted that long-time series are required. It is now possible to lay plans for both geodetic and oceanographic procedures to remedy these deficiencies in the long-term interests of the study.

Revising key parameters for long-term carbon cycle models

2021 ◽  
Author(s):  
Chloé M. Marcilly ◽  
Trond H. Torsvik ◽  
Mathew Domeier ◽  
Dana L. Royer
Keyword(s):  
Sea Level ◽  
Age Distribution ◽  
Silicate Weathering ◽  
Geological Time ◽  
Sea Levels ◽  
Climate Fluctuations ◽  
Climate Gradients ◽  
Sea Level Fluctuations ◽  
Temperature And Precipitation

&lt;p&gt;CO&lt;sub&gt;2&lt;/sub&gt; is the most important greenhouse gas in the Earth&amp;#8217;s atmosphere and has fluctuated considerably over geological time. However, proxies for past CO&lt;sub&gt;2 &lt;/sub&gt;concentrations have large uncertainties and are mostly limited to Devonian and younger times. Consequently, CO&lt;sub&gt;2&lt;/sub&gt; modelling plays a key role in reconstructing past climate fluctuations. Facing the limitations with the current CO&lt;sub&gt;2&lt;/sub&gt; models, we aim to refine two important forcings for CO&lt;sub&gt;2&lt;/sub&gt; levels over the Phanerozoic, namely carbon degassing and silicate weathering.&lt;/p&gt;&lt;p&gt;Silicate weathering and carbonate deposition is widely recognized as a primary sink of carbon on geological timescales and is largely influenced by changes in climate, which in turn is linked to changes in paleogeography. The role of paleogeography on silicate weathering fluxes has been the focus of several studies in recent years. Their aims were mostly to constrain climatic parameters such as temperature and precipitation affecting weathering rates through time. However, constraining the availability of exposed land is crucial in assessing the theoretical amount of weathering on geological time scales. Associated with changes in climatic zones, the fluctuation of sea-level is critical for defining the amount of land exposed to weathering. The current reconstructions used in&lt;sub&gt;&lt;/sub&gt;models tend to overestimate the amount of exposed land to weathering at periods with high sea levels. Through the construction of continental flooding maps, we constrain the effective land area undergoing silicate weathering for the past 520 million years. Our maps not only reflect sea-level fluctuations but also contain climate-sensitive indicators such as coal (since the Early Devonian) and evaporites to evaluate climate gradients and potential weatherablity through time. This is particularly important after the Pangea supercontinent formed but also for some time after its break-up.&lt;/p&gt;&lt;p&gt;Whilst silicate weathering is an important CO&lt;sub&gt;2&lt;/sub&gt; sink, volcanic carbon degassing is a major source but one of the least constrained climate forcing parameters. There is no clear consensus on the history of degassing through geological time as there are no direct proxies for reconstructing carbon degassing, but various proxy methods have been postulated. We propose new estimates of plate tectonic degassing for the Phanerozoic using both subduction flux from full-plate models and zircon age distribution from arcs (arc-activity) as proxies.&lt;/p&gt;&lt;p&gt;The effect of revised modelling parameters for weathering and degassing was tested in the well-known long-term models GEOCARBSULF and COPSE. They revealed the high influence of degassing on CO&lt;sub&gt;2&lt;/sub&gt; levels using those models, highlighting the need for enhanced research in this direction. The use of arc-activity as a proxy for carbon degassing leads to interesting responses in the Mesozoic and brings model estimates closer to CO&lt;sub&gt;2 &lt;/sub&gt;&amp;#160;proxy values. However, from simulations using simultaneously the revised input parameters (i.e weathering and degassing) large model-proxy discrepancies remain and notably for the Triassic and Jurassic.&lt;/p&gt;&lt;p&gt;&amp;#160;&lt;/p&gt;

The Impact of Climate Change on States

2021 ◽  
Vol 23 (2-3) ◽  
pp. 115-132
Author(s):  
Łukasz Kułaga
Keyword(s):  
Climate Change ◽  
International Law ◽  
Sea Level Rise ◽  
Sea Level ◽  
The State ◽  
Fundamental Change ◽  
Sea Levels ◽  
Impact Of Climate Change ◽  
Potential Impact ◽  
The Impact

Abstract The increase in sea levels, as a result of climate change in territorial aspect will have a potential impact on two major issues – maritime zones and land territory. The latter goes into the heart of the theory of the state in international law as it requires us to confront the problem of complete and permanent disappearance of a State territory. When studying these processes, one should take into account the fundamental lack of appropriate precedents and analogies in international law, especially in the context of the extinction of the state, which could be used for guidance in this respect. The article analyses sea level rise impact on baselines and agreed maritime boundaries (in particular taking into account fundamental change of circumstances rule). Furthermore, the issue of submergence of the entire territory of a State is discussed taking into account the presumption of statehood, past examples of extinction of states and the importance of recognition in this respect.

Estimating contribution of high-frequency sea-level oscillations to the extreme sea levels in the Adriatic Sea

2021 ◽  
Author(s):  
Krešimir Ruić ◽  
Jadranka Šepić ◽  
Maja Karlović ◽  
Iva Međugorac
Keyword(s):  
Time Series ◽  
Sea Level ◽  
High Frequency ◽  
Adriatic Sea ◽  
Tide Gauge ◽  
Data Series ◽  
Sea Levels ◽  
Short Period ◽  
Extreme Sea Levels ◽  
Sea Level Oscillations

&lt;p&gt;Extreme sea levels are known to hit the Adriatic Sea and to occasionally cause floods that produce severe material damage. Whereas the contribution of longer-period (T &gt; 2 h) sea-level oscillations to the phenomena has been well researched, the contribution of the shorter period (T &lt;&amp;#160;2 h) oscillations is yet to be determined. With this aim, data of 1-min sampling resolution were collected for 20 tide gauges, 10 located at the Italian (north and west) and 10 at the Croatian (east) Adriatic coast. Analyses were done on time series of 3 to 15 years length, with the latest data coming from 2020, and with longer data series available for the Croatian coast. Sea level data were thoroughly checked, and spurious data were removed.&amp;#160;&lt;/p&gt;&lt;p&gt;For each station, extreme sea levels were defined as events during which sea level surpasses its 99.9 percentile value. The contribution of short-period oscillations to extremes was then estimated from corresponding high-frequency (T &lt; 2 h) series. Additionally, for four Croatian tide gauge stations (Rovinj, Bakar, Split, and Dubrovnik), for period of 1956-2004, extreme sea levels were also determined from the hourly sea level time series, with the contribution of short-period oscillations visually estimated from the original tide gauge charts.&amp;#160;&amp;#160;&lt;/p&gt;&lt;p&gt;Spatial and temporal distribution of contribution of short-period sea-level oscillations to the extreme sea level in the Adriatic were estimated. It was shown that short-period sea-level oscillation can significantly contribute to the overall extremes and should be considered when estimating flooding levels.&amp;#160;&lt;/p&gt;

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