scholarly journals Availability and Structure of Coarse Woody Debris in Hemiboreal Mature to Old-Growth Aspen Stands and Its Implications for Forest Carbon Pool

Forests ◽  
2021 ◽  
Vol 12 (7) ◽  
pp. 901
Author(s):  
Silva Šēnhofa ◽  
Guntars Šņepsts ◽  
Kārlis Bičkovskis ◽  
Ieva Jaunslaviete ◽  
Līga Liepa ◽  
...  

European aspen deadwood is extensively studied as a habitat for saproxylic species, while less is known of its dynamics and role in carbon sequestration. We studied unmanaged mature (41–60 years), moderately overmature (61–80 years), overmature (81–100 years), and old-growth (101–140 years) and managed mature and moderately overmature aspen stands on fertile mineral soils. In unmanaged stands, marginal mean CWD volume was from 67.3 ± 12.1 m3 ha−1 in moderately overmature to 92.4 ± 5.1 m3 ha−1 in old-growth stands, with corresponding marginal mean CWD carbon pool 8.2 ± 1.6 t ha−1 and 12.5 ± 0.7 t ha−1, respectively. Both CWD volume and its carbon pool had substantial yet non-significant differences (all p > 0.05) among the age groups. High CWD volume was present in most stands, by at least two-thirds of plots comprising more than 20 m3 ha−1, and about half of CWD was larger than 30 cm in diameter. Changes in CWD species composition toward a higher proportion of deciduous deadwood in old-growth stands, together with a high volume of recently dead trees, suggest early senescence of the dominant aspen cohort.

2006 ◽  
Vol 36 (10) ◽  
pp. 2614-2622 ◽  
Author(s):  
Tomasz Zielonka

The amount and decay stages of dead wood and the residence time of logs was studied in old-growth spruce-dominated stands in the Tatra and Babia Góra in the western Carpathian Mountains in Central Europe. The DBH of living trees and snags as well as the dimensions of logs (with a minimum diameter of 10 cm) were measured on nine sample plots with a pooled area of 4.3 ha. For the logs, the decay stage was identified according to an eight-level decay classification. Dendrochronological cross-dating was applied to 107 wood samples of logs to determine the time since death. The average volume of living trees was 454 m3/ha and the volume of coarse woody debris (CWD) averaged 191 m3/ha. The noted volume of CWD was higher than volumes reported from previous studies in the Carpathian Mountains. On average, CWD made up 30% of the total volume (living plus dead) and varied between 49% and 21% for the stands. The average age of logs for decay class 1 was 13 years and 24, 28, 38, 45, 38, and 60 years for decay classes 2–7, respectively. Considerable variation of time since death between decay classes was probably due to a variable time while dead trees remained standing as snags.


2015 ◽  
Vol 76 (4) ◽  
pp. 322-330 ◽  
Author(s):  
Konrad Skwarek ◽  
Szymon Bijak

Abstract Dead wood plays an important role for the biodiversity of forest ecosystems and influences their proper development. This study assessed the amount of coarse woody debris in municipal forests in Warsaw (central Poland). Based on the forest site type, dominant tree species and age class, we stratified all complexes of the Warsaw urban forests in order to allocate 55 sample plots. For these plots, we determined the volume of dead wood including standing dead trees, coarse woody debris and broken branches as well as uprooted trees. We calculated the amount of dead wood in the distinguished site-species-age layers and for individual complexes. The volume of dead matter in municipal forests in Warsaw amounted to 38,761 m3, i.e. 13.7 m3/ha. The obtained results correspond to the current regulations concerning the amount of dead organic matter to be left in forests. Only in the Las Bielański complex (northern Warsaw) volume of dead wood is comparable to the level observed in Polish national parks or nature reserves, which is still far lower than the values found for natural forests. In general, municipal forests in Warsaw stand out positively in terms of dead wood quantity and a high degree of variation in the forms and dimensions of dead wood.


1994 ◽  
Vol 24 (10) ◽  
pp. 1989-1996 ◽  
Author(s):  
Glenn H. Stewart ◽  
Larry E. Burrows

The volume, biomass, and carbon and nitrogen content of coarse woody debris were measured on three 1-ha reference plots in old-growth Nothofagusfusca (Hook. f.) Oerst.–Nothofagusmenziesii (Hook. f.) Oerst. forest on the South Island of New Zealand. Two decay sequences for logs and one for standing dead trees (snags) were recognised from two-way indicator species analysis (TWINSPAN) of up to 30 variables related to physical characteristics and structural integrity. Wood volume (up to 800 m3•ha−1) and biomass were high (up to 300 Mg•ha−1), and the inside-out decay sequence from heartwood to sapwood was unusual compared with that of other temperate hardwood forests. Coarse woody debris represented significant carbon and nitrogen pools, with ca. 150 Mg•ha−1 and 370 kg•ha−1, respectively, in one stand. The coarse woody debris component of these broad-leaved evergreen hardwood forests was much higher than that reported for other temperate hardwood forests and approaches that of many northern hemisphere conifer forests. The large coarse woody debris pools are discussed in relation to live stand biomass, natural disturbances and tree mortality, and decomposition processes.


1990 ◽  
Vol 20 (3) ◽  
pp. 326-330 ◽  
Author(s):  
M. H. McDade ◽  
F. J. Swanson ◽  
W. A. McKee ◽  
J. F. Franklin ◽  
J. Van Sickle

Coarse woody debris from streamside forests plays important biological and physical roles in stream ecosystems. The distance from stream bank to rooting site was determined for at least 30 fallen trees at each study site on 39 streams in the Cascade and Coast ranges of Oregon and Washington. The study sites varied in channel size (first- through third-order), side-slope steepness (3 to 40°), and age of surrounding forest (mature or old-growth stands). The distribution of distances from rooting site to bank was similar among streams, with 11% of the total number of debris pieces originating within 1 m of the channel and over 70% originating within 20 m. Stands with taller trees (old-growth conifers) contributed coarse woody debris to streams from greater distances than did stands with shorter (mature) trees.


2010 ◽  
Vol 26 (4) ◽  
pp. 467-471 ◽  
Author(s):  
Lisa B. Kissing ◽  
Jennifer S. Powers

The ecological importance of trees lasts much longer than their life spans. Standing dead trees (snags) and fallen trunks and branches are an important component of above-ground carbon stocks and nutrient reserves, provide habitat for wildlife, and interact with disturbance regimes (e.g. by serving as fuel for fires) (Clark et al. 2002, Harmon et al. 1986, Pyle et al. 2008). Despite these diverse functions, woody debris stocks remain poorly quantified in tropical forests in general (Brown 1997), and in tropical dry forests in particular (Harmon et al. 1995). More empirical studies of the patterns of woody debris and processes that control its dynamics are needed to understand its role in global biogeochemical cycles and for ecosystem simulation models, many of which do not represent coarse woody debris (CWD) as a separate pool (Cornwell et al. 2009).


2003 ◽  
Vol 11 (S1) ◽  
pp. S135-S157 ◽  
Author(s):  
M C Feller

This paper synthesizes data extracted from the literature and data collected in various studies by the author on the quantity, characteristics, and functional importance of coarse woody debris (CWD) in the old-growth forests of British Columbia (B.C.). There is little agreement in the literature about the minimum diameter of CWD or the number of decay classes recognized. In western North America, five decay classes are commonly used, but recent studies suggest fewer decay classes are preferable. Comparisons among decay classes and biogeoclimatic zones and subzones in B.C. reveal that quantities and volumes are greatest (up to approximately 60 kg/m2 and approximately 1800 m3/ha, respectively), and CWD persists the longest (sometimes in excess of 1000 years) in the Coastal Western Hemlock (CWH) biogeoclimatic zone. The quantity and ground cover of CWD increase with forest productivity. Persistence of CWD has varied from less than 100 to over 800 years in two coastal (CWH and Mountain Hemlock (MH)) and three interior (Interior Douglas-fir (IDF), Interior Cedar–Hemlock (ICH), and Engelmann Spruce – Subalpine Fir (ESSF)) biogeoclimatic zones. Trends in CWD quantity with forest age in managed coastal B.C. forests suggest a U-shaped curve, with greater quantities occurring in recent cutovers than in old-growth forests, and lowest quantities occurring in middle-aged forests. This may be the normal trend in CWD with forest age, with departures from this trend resulting from disturbance- or environment-specific factors. Relatively large amounts of data exist on the characteristics of CWD in the CWH, IDF, ICH, ESSF, and Boreal White and Black Spruce (BWBS) biogeoclimatic zones, but such data for the Coastal Douglas-fir, Sub-Boreal Pine–Spruce, Sub-Boreal Spruce (SBS), and Spruce–Willow–Birch biogeoclimatic zones appear relatively sparse. There have been few studies of the functional role of CWD in B.C. forests, but those studies that have been completed indicate that CWD is an important habitat component for some plant and animal species. A total of 169 plant species, including >95% of all lichens and liverworts, were found to grow on CWD in old-growth forests in the CWH, MH, IDF, ICH, and ESSF biogeoclimatic zones. One third of these species were restricted to CWD. Studies in several biogeoclimatic zones have found that CWD provided preferred habitat for and was associated with higher populations of some small animal species, such as shrews, some voles, and some salamanders, in old-growth forests, but the effects varied with species and biogeoclimatic zone. The nutrient cycling role of CWD is not yet well known, but it currently appears to be relatively insignificant in B.C. old-growth forests. Although it has been considered that CWD could increase mineral soil acidification and eluviation, no evidence for this was found in a study of the CWH, MH, IDF, ICH, ESSF, BWBS, and SBS biogeoclimatic zones. Future studies of the functional role of CWD should consider both scale (square metre vs. hectare) and temporal (changes in CWD with forest age) issues, as studies including these are sparse and both may be important. Key words: biogeoclimatic zones, British Columbia, coarse woody debris, old-growth forests.


2016 ◽  
Vol 12 (9) ◽  
pp. 69
Author(s):  
Carlos Belezaca Pinargote ◽  
Darwin Salvatierra Piloso ◽  
Diana Delgado Campusano ◽  
Roberto Godoy Bórquez ◽  
Eduardo Valenzuela Flores ◽  
...  

Coarse woody debris (CWD) are the necromass in wooded environments and comply with various ecosystems functions, such as seedling nursery, habitat other organisms, store carbon (C) and nutrients, etc.. Volume, necromass, and decay states of CWD dead in an old-growth temperate (OGTF) forest in Puyehue National Park, South-Central Chile were evaluated. In 10 plots of 900 m2 CWD was quantified (≥ 10 cm diameter), whose necromass classified using a scale of five categories/stages of decay, necromass (1 = lowest and 5 = highest degradation). The average forest density was 299 trees ha-1, 112 m2 ha-1 of basal area, and 2.395 m3 of stem volume. The upper arboreal stratum was dominated by N. betuloides. The greatest amount of CWD belonged to N. betuloides (95,2%), where logs (52,7%) and branches (35%), plant structures were present in greater numbers. The bulk of necromass found in advanced states of decomposition (level 4 and 5) with 56,5% and 34,1%, respectively. It was determined that the CWD volume was 632 m3 ha-1, representing a necromass of 321.5 Mg ha- 1. These results demonstrate that the old-growth temperate forests of southern Chile are significant reserves of coarse woody debris, which contributes to the biogeochemistry of these complex and remote ecosystems.


2021 ◽  
Vol 4 ◽  
Author(s):  
Jeffrey Opoku-Nyame ◽  
Alain Leduc ◽  
Nicole J. Fenton

Clear cut harvest simplifies and eliminates old growth forest structure, negatively impacting biodiversity. Partial cut harvest has been hypothesized (1) to have less impact on biodiversity than clear cut harvest, and (2) to encourage old growth forest structures. Long-term studies are required to test this hypothesis as most studies are conducted soon after harvest. Using epixylic bryophytes as indicators, this study addresses this knowledge gap. Fourteen years after harvest, we examined changes in epixylic bryophyte community composition richness and traits, and their microhabitats (coarse woody debris characteristics and microclimate) along an unharvested, partial cuts and clear cuts harvest treatment in 30 permanent plots established in the boreal black spruce (Picea mariana) forests of northwestern Quebec, Canada. Our results were compared to those of an initial post-harvest study (year 5) and to a chronosequence of old growth forests to examine species changes over time and the similarity of bryophyte communities in partial cut and old growth forests. Coarse woody debris (CWD) volume by decay class varied among harvest treatments with partial cuts and clear cuts recording lower volumes of early decay CWD. The epixylic community was richer in partial cuts than in mature unharvested forests and clear cuts. In addition, species richness and overall abundance doubled in partial and clear cuts between years 5 and 14. Species composition also differed among treatments between years 5 and 14. Furthermore, conditions in partial cut stands supported small, drought sensitive, and old growth confined species that are threatened by conditions in clear cut stands. Lastly, over time, species composition in partial cuts became more similar to old growth forests. Partial cuts reduced harvest impacts by continuing to provide favorable microhabitat conditions that support epixylic bryophytes. Also, partial cut harvest has the potential to encourage old growth species assemblages, which has been a major concern for biodiversity conservation in managed forest landscapes. Our findings support the promotion of partial cut harvest as an effective strategy to achieve species and habitat conservation goals.


2010 ◽  
Vol 25 (4) ◽  
pp. 176-180 ◽  
Author(s):  
David Azuma

Abstract Forest Inventory and Analysis data were used to investigate the effects of a severe western spruce budworm outbreak on the dead wood component of forests in 11 counties of eastern Oregon for two time periods. The ownership and the level of damage (as assessed by aerial surveys) affected the resulting down woody material and standing dead trees. The pattern of coarse woody debris with respect to ownership and management intensity remained consistent into the next 10-year period. Harvesting tended to lower the amount of coarse woody debris on private forests. Federally managed forests had more standing dead trees than private lands, with more in the reserved than nonreserved areas. There was a reduction in the number of standing dead trees between the two periods.


2019 ◽  
Vol 65 (No. 10) ◽  
pp. 408-422
Author(s):  
Vahid Etemad ◽  
Mohsen Javanmiri pour ◽  
Zeinab Foladi

In a natural forest, phases of different dynamics are gradually replaced to create sustainability in the stands. Coarse woody debris is among the most significant structural elements of natural stands that perform an influential position in the identification of dynamic phases. Therefore, the focus of this study is on dead wood conditioning as one of the major structural components in determining the various dynamic phases in the northern forests of Iran as part of the temperate forests. For this study, compartment 326 of Gorazbon District was considered as one of the control parcels of Kheyroud Forest. In this parcel, 25 one-hectare sample plots were selected as permanent plots for a long-term forest structure and succession studies. The coarse woody debris by 100% sampling method was measured. The results showed that there are 8 main phases in this area (gap formation, understorey initiation, stem exclusion, volume accumulation, volume degradation, multiple, lighting, old-growth). The extensive forest area (52%) is located in the understorey initiation and stem exclusion phases. The results also showed that the total average volume of snags and logs was 41.5 m<sup>3</sup>·ha<sup>–1</sup>. Furthermore, the mean dead wood volume in decay classes 1, 2, 3 and 4 was 10.33, 12.22, 9.15 and 83.9 m<sup>3</sup>·ha<sup>–1</sup>, respectively. The average frequency of dead trees in the diameter classes smaller than 25 cm, 25–50 cm and in the diameter class more than 50 cm is 25.79, 6.93, and 4.88. The significance analysis results obtained by ANOVA test showed that there is a significant difference between volume, snag and log stock and the shape of dead wood in various dynamic phases. Therefore, in general, dead wood in the forest differs according to habitat, evolutionary stage (dynamic phases), standing volume and species diversity of the tree species.


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